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This scientific review analyzes 191 field studies published since 1995 in order to identify beekeeping practices and biosecurity measures that have demonstrated a measurable effect on colony health and productivity. In total, 744 “practice records” were extracted and structured according to theme, region, and season of application. The objective is to propose an analytical framework to guide beekeeping decisions based on comparative trials conducted under real-world conditions. The methodically structured review character is central to this synthesis, which highlights both dominant practices and research gaps.
Swarming is the bee colony’s natural mode of reproduction. In a prime swarm, the mated old queen leaves the nest with a large part of the adult workers, while the mother colony retains the combs, the brood, the food stores and one or more young queens in development. The division is strongly asymmetric: a direct measurement of naturally swarming colonies showed that roughly three quarters of the adult workers could leave on average with the prime swarm, with considerable variation between colonies (Rangel & Seeley, 2012).
For the beekeeper, however, the essential question is not only whether a colony is likely to swarm, but where it stands within the process. An intervention that is useful several weeks before departure may become insufficient once queen rearing is well under way.
Four types of queen cells are distinguished :
1. Natural swarming queen cells
2. Emergency queen cells
3. Supersedure queen cells
4. Artificial queen cells
The amateur beekeeper is often well equipped to manage their colonies: observant, curious, motivated, enthusiastic, sometimes idealistic, often well coached, supported by introductory courses and the reading of numerous articles online; they share their concerns with colleagues and appreciate advice or even hands-on help from the “experienced” beekeepers… yet they often have no clear idea of what their hobby actually costs or earns! Reassurance comes from weighing the harvest: great, 15 kg of honey per colony!!! But what did those 30 jars they are so proud of really cost? What is the exact profit from selling the part of the production that was not consumed personally?
The experienced beekeeper asks a different question: what if I turned this hobby into my profession? What is the initial investment, and what turnover do I need to achieve to make a living?
The professional beekeeper must report to their bank and asks: how can I improve the profitability of my operation? How can I depreciate assets in order to renew my equipment?
How can risks related to poor harvest years or diseases be managed?
All three types of beekeepers have access to a user-friendly tool, developed with expertise and precision, to help them answer all questions related to financially responsible beekeeping.
Good beekeeping practice encompasses the fundamental principles for ensuring responsible, sustainable beekeeping that respects bee colonies. It provides a reference framework for all beekeeping activities. The information below is based on summary guide 4 from the Swiss Bee Health Service.

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Water is indispensable to a honey bee colony, but its needs vary considerably with temperature, brood rearing, food resources and season. From the water balance of individual bees to colony thermoregulation, the dilution of larval food and the concentration of nectar, this article explains the many flows of water within the hive. It derives practical guidance for water provision, periods of heat and overwintering.

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What is the legal status of beekeepers in Switzerland? When does a beehive become a structure requiring a permit? This article summarizes the legal opinion drafted in 2023 by Prof. Thierry Largey (University of Lausanne), whose conclusions shed light on beekeeping practices without, however, replacing a legal analysis of a specific case.
Diarrhoea in bees is most often caused by Nosema or dysentery. Nosema is a fungal disease that can be caused by two different pathogens: Nosema apis and Nosema ceranae. Transmission occurs in the form of spores via food, water, and contaminated surfaces. The fungus germinates in the midgut and subsequently multiplies in the intestinal cells. New spores are transmitted through the excreta. In this factsheet, only diarrhoea resulting from Nosema apis is addressed and discussed. The disease is strongly season-dependent and occurs mainly in spring. The pathogen can be detected within the colony throughout the year even in the absence of clinical symptoms.
October marks the transition to wintering: major tasks are complete, but a few checks remain crucial. These mainly involve verifying food stores, colony strength, varroa mite infestation, and apiary protection, and then minimizing disturbances. The goal is simple: to enter winter with healthy, compact, and well-prepared colonies.
The mānuka (Leptospermum scoparium), also called the New Zealand tea tree in English because its leaves can be used to make tea, is an evergreen perennial shrub belonging to the family Myrtaceae (the same family as eucalyptus and myrtle). It has irregularly branched shoots, and its leaves are small, variably shaped, and aromatic. During flowering, mānuka produces five-petaled flowers that may be white, pink, or red, followed by fruits in the form of round, woody capsules that persist on the plant for many months.
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Wax combs age together with the colony: they darken, accumulate cocoons, lose cell volume and may store residues. This article summarises what recent studies show about the effects of old combs on brood, bees and the quality of bee products, and draws practical recommendations for the apiary.

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Why does a colony replace a newly introduced queen within weeks, yet keep another despite a patchy brood pattern? This article explains how signals from the queen, brood and colony interact, and what this means for assessing, introducing and caging queens.
Cape ragwort (Senecio inaequidens) is a perennial herbaceous plant belonging to the Asteraceae (or Compositae) family, native to South Africa, from where it was accidentally introduced through the wool trade. Its branched stem reaches a height of 40 to 60 cm and bears yellow flowers that bloom over a long period, from early summer through autumn, attracting bees mainly for their yellow pollen.
The queen has a lifespan of 4–5 years. This may seem long. However, it takes 1–2 years to identify a valuable queen and select her for (re)production. To preserve this queen for as long as possible, her egg laying must be managed carefully, as it depends on the number of spermatozoa present in her spermatheca. Various techniques make it possible to modulate egg laying: colony size, the surface of empty cells, the population of young bees, protein resources …
One might assume that asexual reproduction leads to an evolutionary dead end, with the accumulation of deleterious mutations ultimately resulting in species extinction. Recent studies show that this is not the case, and the drone provides a well-studied example. The particular case of reproduction in Varroa destructor is illuminating: these mites exhibit fewer deleterious mutations than other sexually reproducing insects …
Examples of asexual reproduction in certain animals include: the hammerhead shark, the leopard shark, the sawfish, the sea anemone, the Komodo dragon, the gecko, the aphid, the scale insect …
Practical Guide 4.7.5 presents spring swarming as a natural behaviour essential to the survival of the bee population. It also notes that swarms that depart have little chance of surviving long-term in the wild in Switzerland, that they can transmit diseases, and that they can cause conflicts or dangerous situations in urban areas. For beekeeping, the departure of a swarm generally means a loss of honey yield and requires particular attention to the colony that has swarmed.
Recognizing healthy colonies is essential for effective beekeeping and for early detection of potential problems. The official checklist 4.7.3 from the Swiss Bee Health Service describes the typical characteristics of healthy colonies and provides reliable observation guidelines.
This review article analyses honeybee colony losses from an immunological perspective, focusing on how diverse environmental and ecological stressors affect immune competence. It synthesizes current knowledge on the innate and social immune mechanisms of Apis mellifera and examines how pathogens, parasites, pesticides and nutritional stress interact. Particular attention is given to synergistic effects between stressors and their implications for colony collapse disorder. The review also discusses nutritional, natural-product and technological strategies proposed to support honeybee health, while highlighting the need for integrated and evidence-based management approaches.
Queen rearing is one of the fundamental pillars of modern beekeeping. The quality of a colony depends largely on its queen. Through her genetic heritage and constant production of pheromones, she influences the vitality, behavior, and performance of the entire colony.
Controlled breeding is not limited to the production of new queens. It is based on a thorough understanding of biological mechanisms—swarming tendency, orphanhood, fertilization—as well as solid genetic foundations and structured selection.
The European honey bee, also known as the honey fly (Apis mellifera), is a domesticated honey bee species native to Europe. It is considered semi-domesticated. It is one of the bee species bred on a large scale for honey production.
There are several circumstances in which this beekeeping operation is required; however, the most serious case, which calls for the greatest precautions, is American foulbrood. The content of this information sheet therefore focuses on this specific disease.
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To successfully raise queens, it’s not enough to produce high-quality queen cells: the quality of the drones is just as important. This article summarizes what we know about their development, sexual maturity, the impact of Varroa mites, and the limitations of controlling fertilization in open-hive systems.
The honey bee is the fifth insect and the first hymenopteran whose genome has been fully sequenced. The DNA analysis was based on males originating from a single queen obtained from the Bee Weaver apiaries in Texas. Since the project began, approximately 14 million individual reads were required to reconstruct the 236 million base pairs that constitute the nearly complete honey bee genome.
Hygiene in colony management is essential for preventing disease and maintaining healthy colonies. It concerns both the beekeeper's behavior and the correct use of equipment. The following information is based on the official checklist 4.1 of the Swiss Bee Health Service.
The flight hole protected by a grid is a protective measure designed to reduce predation by Asian hornets in front of colonies. This adaptation allows bees to move around normally while limiting access to predators. The following information is based on the official checklist 2.7.1 of the Swiss Bee Health Service.
A queen or artificial swarm is a method for establishing a new colony with a defined queen. It is deliberately created to build strong, vigorous colonies, form nucleus colonies, or carry out rehabilitation measures. The summary below is based on the official Practical Guide 1.4.3 of the Swiss Bee Health Service.
The small hive beetle (Aethina tumida) is a feared pest of honey bee colonies; both adults and larvae feed on honey, pollen, and preferably brood. European countries had so far been considered free of the small hive beetle. Reports from September 2014 concerning the detection of Aethina tumida in southern Italy are alarming and also require increased vigilance from Swiss beekeepers, as well as regular inspection of their own bee colonies.
The queen’s primary function is to lay eggs, enabling the emergence of all individuals that make up the population of a bee colony. The queen’s many other functions are not addressed in this brief article.
The weakening of a bee colony is rarely the result of a single factor: it is most often a gradual process in which several constraints interact and reduce the resilience of the superorganism. This summary presents a conceptual model organized into three main self-amplifying spirals: an infection spiral, a cooling spiral, and a starvation spiral. As long as the population remains sufficient and healthy, the colony can compensate for temporary disturbances, but when resilience declines, regulatory mechanisms become fragile and the dynamics of weakening accelerate. The practical benefit is to help beekeepers recognize weak signals earlier and intervene before several spirals set in simultaneously.
A hive inspection should be prepared in advance to avoid omissions and to improve efficiency. Any inspection, even a brief one, disturbs the colony, which perceives it as an intrusion; the resulting stress causes the bees to consume between 0.5 and 1 kg of honey. Inspections should therefore only be carried out when necessary. The following tips make inspections more effective:
In 2019, it came as a surprise to learn that the Varroa mite does not feed on the bees’ haemolymph but pierces their cuticle in order to ingest the so-called fat body (► see article).
Research is progressing very rapidly, and today it is known that the saliva of the adult Varroa mite allows it to keep the opening it has pierced through the membrane of the nymph or the imago wide open, in order to liquefy and then suck up the fat body. A salivary enzyme (chitinase) appears to reduce the bee’s immune defences and to play an important role in the survival of the Varroa mite … it is therefore natural to envisage blocking this enzyme as a future strategy in the fight against this deadly parasite …
In beekeeping supply stores, frames are available pre-assembled with or without wax, as well as frames to be assembled. For a pre-assembled frame with wax, a minimum cost of CHF 10 should be expected.
Measuring about 1 mm³, the bee’s brain is capable of remarkable feats thanks to highly sophisticated cognitive mechanisms. The bee makes decisions, memorises flight paths by precisely tracking the course of the sun, identifies food sources and evaluates them both qualitatively and quantitatively, and transmits multiple types of information to its nestmates in order to recruit a large number of foragers with differentiated skills.
A brood break followed by oxalic acid treatment can be an alternative to the first summer formic acid treatment against varroa. This article explains when this approach may be useful, which biological conditions must be met, and which checks are essential for safe and effective use.
What makes a good queen: her origin, the rearing method or the type of mating? This article explains why terms such as natural, grafted, F1, F2 or artificial insemination do not, on their own, determine queen quality. What matters is the whole chain: suitable starting material, careful rearing, an appropriate mating strategy and the later performance of the colony.
The many scientific articles available to beekeepers provide valuable insight into the physiology of honey bees, drones and, of course, Varroa destructor. What is crucial, however, is to understand the dynamic relationships between these different actors within the colony over time—that is, the combined pathophysiology of these three populations throughout the season.
Wintering is a crucial phase in the beekeeping year and largely determines the development of colonies in the following spring. It depends on colonies that are healthy, sufficiently strong, and properly prepared. The information below is based on the official BGD/SSA checklist 4.3 and summarizes the key points for successful wintering.

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Beekeeping in Switzerland is not regulated by a single law, but is subject to several areas of law, in particular animal disease law and—in the case of honey sales—food law, as well as other regulations such as animal welfare or building and environmental law. This article presents the core obligations under federal law in a practical and legally precise manner and clarifies questions regarding registration, documentation, disease outbreaks, and sales. A concluding checklist enables systematic self-assessment of legal compliance.
Matthieu Guichard (Agroscope, Swiss Bee Research Centre, Bern) conducted a long-term study between 2010 and 2018 on approximately 1000 Carnica honey bee colonies and approximately 1000 Mellifera honey bee colonies, aiming to determine the heritability of various traits: honey yield, gentleness, comb adherence, swarming tendency, hygienic behaviour, and Varroa infestation.
To unravel the mysteries of crystallization, the authors explore the world of science and delve into the heart of the product: principles of chemistry and physics, together with a measure of common sense, are essential to understand it and to highlight the different factors that can influence it.
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Queen ringing is presented as an emerging method to temporarily interrupt egg laying without confining the queen. This summary examines what the first studies actually show, what remains uncertain, and what Swiss beekeepers can — or cannot yet — cautiously take from it for Varroa control.
The bee dance is a term used in apiculture and ethology to describe a system of animal communication by which foraging or scouting bees (on average 5 to 25 % of the foragers, which are the oldest and most experienced; the others act as receivers waiting for the scout’s signal) convey to the receivers remaining in the colony the distance and direction of the food source where they can obtain the nectar and pollen from flowers required for honey production.
The colour of pollen varies depending on the plant species: it ranges from white to black, but most often pollen is yellow or orange, as well as light brown or greenish. Based on colour, certain typical pollens can be recognised (for example, poppy pollen is black and phacelia pollen is blue), but only microscopic analysis allows precise identification of the pollen type.
The ITSAP (http://blog-itsap.fr/) participated in a European trial on the brood interruption technique in varroa control, which aimed to quantify efficacy, the impact on colony development, and the practicality of different approaches, whether combined or not with the use of oxalic acid. Here is a summary of the results and discussion of the scientific article (Büchler et al.) published this year in the Journal of Apicultural Research.
Metamorphosis is a major transformation of the body and way of life during the development and life cycle of certain animals, such as amphibians and some insects. It represents an evolutionary adaptation of an organism to its respective environmental conditions. In zoology, it refers to the transformation from the larval form to the adult, sexually mature animal.
by ALAIN SATABIN
As early as Antiquity, mathematicians observed that the shape of honeycomb cells in beehives optimizes the ratio between available volume and the amount of wax used. Demonstrating this, however, is another matter … one that has still not been fully resolved.
Impact of varroa infestation on thermoregulation in honey bee colonies: new findings indicate that varroa mites reduce bees’ ability to effectively regulate their body temperature, making them more vulnerable to cold temperatures.

Cantharis fusca | Flug | Garten | Gemeiner Weichkäfer | Insektensterben | Käfer | artensterben | bestäuben | bestäubung | blütenpflanzen | gifte | glyphosat | gärten | pestizide | roundup | steingärten | tot | umweltverschmutzung | wahnsinn | wildbienen
© Lothar Lenz
Pollinators are much more than honey bees. This article provides a scientifically grounded and practical overview of their diversity, their role in ecosystems and agriculture, the main pressures they face, and concrete measures to support them.
In the hive, as we have seen, water serves multiple purposes: it is indispensable for both mature and immature individuals and also enables bees to regulate the nest climate during periods of high heat. As water is hardly stored within the hive, it must be brought in from outside as needs arise. This task is carried out by water carriers, specialised foragers that continue their work even under adverse conditions.
(by Janine Kievits)
A drone-laying colony normally has no future. It can raise only males, either because its queen has lost the ability to fertilise the eggs she lays, or because there is no queen at all and laying workers have taken over. And yet, from time to time—very rarely, but it does happen—a beekeeper is surprised to discover a fine brood developing in a hive whose frames he was about to shake out. How can this be possible?
A brood interruption is a deliberate suspension of brood production in honey bee colonies, used as a measure in Varroa management (e.g. for Varroa reduction or preparation for treatments). The following summary is based on the official Practical Guide 1.6.1 of the Swiss Bee Health Service.
What qualities are required to be a good beekeeper? They are numerous, as beekeeping draws on a wide range of techniques and fields. Some are present from the outset, while others are acquired over time.
Orphan colonies are a special case in beekeeping, as the absence of a queen prevents reproduction and compromises the colony's survival. Official memo 4.7.4 from the Swiss Bee Health Service describes the possible causes of orphanhood and the appropriate measures for managing these colonies.
The deformed wing disease virus (DWV) is one of the most widespread viruses in Apis mellifera and one of the least virulent. In the absence of facilitating factors, it causes only a covert infection, that is, an infection without clinical signs.
The adaptive strategies of imitation make mimicry a model of a complex coevolutionary mechanism involving a first species acting as a model (e.g. Apis cerana) and a second, imitating species (e.g. Varroa destructor), very often parasitic. Varroa destructor, a parasitic mite of bees, has the ability to mimic the chemical composition of its host’s cuticle; moreover, it is also capable of modifying this composition according to the species it parasitizes.
The artificial swarm is a beekeeping method for establishing new colonies in which bees are deliberately taken from existing colonies and reassembled. It serves, among other purposes, colony multiplication, rehabilitation of affected colonies, and reduction of varroa. The following information is based on the official Practical Guide 1.4.2 of the Swiss Bee Health Service.
Periods of scarcity are times when bees cannot find sufficient sources of nectar and pollen, which can lead to food shortages for the colony. The summary below is based on official guidance document 3.2 from the Swiss Beekeeping Service (BGD/SSA) and outlines the causes, effects, and practical management measures.
American foulbrood is a serious and highly contagious brood disease. Incorrectly regarded by beekeepers as a shameful disease, many preconceived ideas about it persist. This is a good opportunity to revisit and clarify the subject.
The search for a suitable apiary location is subject to several conditions, including access to nectar and pollen resources. It is therefore important to be able to identify the different local plant species in order to assess the availability of these resources, or even to increase them if possible.
The terms F0, F1, F2 or Fx are often used in queen breeding, but they do not by themselves indicate whether a queen is good or poor. This article explains what these terms mean, what role heterosis may play, and why decisions should always be based on origin, mating and, above all, the real performance of the colony.
A honey bee colony cannot exist without a queen. She is the only fertile female individual in the colony. Normally, there is only one mated adult queen per hive. She is usually the mother of most, if not all, of the bees in the hive. Through the pheromones she secretes, the queen is able to influence the bees of her colony. In addition, characteristics such as vitality, gentleness, swarming behaviour, disease resistance, and performance are largely determined by the queen.
Regular inspection of colonies makes it possible to identify potential health problems at an early stage and to implement planned and targeted measures. It is essential to check and assess colonies at least in spring (when grape hyacinths are in bloom), in summer (between the summer honey harvest and the first summer treatment), and in October (before overwintering).
Numerous articles available on the internet provide valuable advice on how to manage our bee colonies. It can sometimes be useful to return to more basic notions, grounded in common sense. The “ten mistakes to avoid” are intended as a way to spark the curiosity of beginner beekeepers or to encourage reflection among more experienced colleagues.
Fortunately, there are not only ten mistakes to avoid… otherwise beekeeping would be tedious or even wearisome!
Planning the beekeeping season makes it possible to prepare optimally for upcoming activities with our bees. Thanks to the operating concept developed by apiservice, this is easy to implement for all beekeepers. It is an indispensable tool for effective apiary management. The operating concept can be personalised and allows for a clear visualisation of beekeeping activities and the planning of the appropriate work.

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Pollen is essential for brood rearing and for the turnover of the worker population. Its role cannot, however, be reduced either to a protein percentage or to the number of pellets observed at the hive entrance. To assess a colony's supply, the available quantity, the composition of the pollens, their complementarity and the continuity of income must be considered together.
Official Practical Guide 2.12 classifies Tropilaelaps as a monitored disease. It describes two species of parasitic mites originating from Asia that can infest the European honey bee, and lists patchy brood, sunken/perforated cappings, and deformed bees as typical infestation indicators. For Switzerland, the document states that no presence has been confirmed to date, and that refraining from bee imports, conducting regular monitoring, and notifying the bee inspector in the event of suspected infestation are key preventive measures.

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The winter months can be used to clean all beekeeping equipment. This may not only help to prevent certain diseases, but it is also a pleasure to find tools in good condition in spring, and above all tools that are not sticky.
Every May 20, World Bee Day returns to the calendar. Behind this date lies a broader question: Why have we dedicated a day to pollinators? What is its specific purpose, and what does science tell us about the actual status of these species? The following provides some context based on the available evidence (See the official website: World Bee Day).
The Institute of Apicultural Research at the University of Hohenheim has discovered an active substance against varroa: lithium chloride.
This is very good news, although the active substance—before being placed on the market as a medicinal product for bees—must of course undergo further testing to determine the optimal dosage and to rule out side effects for bees and users as well as the risk of residues. It will therefore still take some time before varroa control can be optimised.
Colony multiplication based on swarming impulse makes targeted use of the natural reproductive drive of honey bee colonies for the formation of nucleus colonies. When correctly applied, this method allows simple multiplication and the use of existing queen cells, but it requires careful observation and timely intervention. The following information is based on the official Practical Guide 1.4.7 of the Swiss Bee Health Service (BGD).
As will be clear, effective control of varroa relies on anticipation rather than on reaction alone. The objective is to keep infestation levels low throughout the season in order to preserve the colony and prevent a high parasitic and viral load. Particular emphasis is placed on protecting the winter bees so as to ensure that the colony can restart under the best possible conditions. Above all, the strategy must be adapted to the season in progress. Each year is unique, and climatic variations require increasing adaptability in order to keep varroa under control.

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Infestation-oriented Varroa treatment (July to September) deviates, according to official Practical Guide 1.6.6, from the Varroa concept 1.1 and is intended for beekeepers who wish to treat only when necessary. It requires monthly, careful assessment of natural mite fall and individual evaluation of each colony. The following summary is based exclusively on the official Practical Guide (BGD/SSA).
The beekeeping calendar presented below was created by a hobby beekeeper using his own approach. The tasks to be carried out each month depend strongly on the region, altitude, type of hive, type of bees, and many other factors. This calendar was designed to help beginners in our association and our region manage their first hives throughout the year and to provide answers to the questions they may have from month to month.
These images are royalty-free and available for free download.
Please credit the author: S. Imboden – www.ApiSavoir.ch
Bee poisoning can cause significant colony losses and requires a rapid and structured response. This checklist describes the characteristic signs, possible causes, and steps to take if poisoning is suspected. The following information is based on the official checklist 3.1.2 of the Swiss Bee Health Service.
Every beekeeper is required to indicate the following information on each jar of honey sold (ODAIOUs Art. 26, food labelling ordinance, OEDAI):
Ellingsenius fulleri is the most frequently encountered pseudoscorpion species of the family Cheliferidae in South Africa. This arthropod is considered a predator of small mites and wax moth larvae found in debris at the bottom of beehives. They often attach themselves to the legs of bees and thus appear to spread to other colonies.
Bee losses during mowing can be significant, particularly on flowering areas that are heavily frequented by foraging bees. Official memo 3.3 from the Swiss Bee Health Service describes the causes of these losses and measures to reduce them through appropriate mowing practices.
Reviewed for you by Claude Pfefferlé
But where does the queen fly? The unprecedented case of a foraging queen! Another dogma is being challenged…
In the countryside of northern Sardinia, an Italian honey bee queen (Apis mellifera ligustica) was observed for the first time in spring 2021 while foraging on a borage flower (Borago officinalis), most likely during an orientation flight prior to mating.
This article provides a structured synthesis of current scientific knowledge on the factors influencing honey quality. It is based exclusively on a recent literature review, examining honey composition, quality criteria, the impact of beekeeping and environmental practices, as well as the links between colony management, honey processing, and biological properties.
The Apidea dispenser is an approved evaporation system for the application of formic acid in small mating units and nucleus colonies. Due to the high sensitivity of these units, strict adherence to the prescribed procedure is particularly important. The following information is based exclusively on the official Practical Guide 1.2.4 of the Swiss Bee Health Service.

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© Lothar Lenz lotharlenz@gmx.de
This article provides an accessible overview of the main issues surrounding wild bees in Switzerland. For a more in-depth discussion, with greater scientific detail, references, and further analysis of ecology, threats, and conservation measures, please refer to the main feature article. This condensed version summarizes the key points in a shorter, more practical format.
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This study shows that water collection increases with colony size and on warm days. It also serves as a reminder that water is a basic resource for brood and the normal functioning of the colony. For the apiary, the message is straightforward: a reliable watering point is worth planning for, with appropriate caution given the semi-controlled experimental setup.

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The bee’s gut microbiota plays an important role in digestion, pollen utilization, and certain defenses against pathogens. This overview also shows that feeding is not merely a matter of providing sugars or proteins, and that natural pollen remains the biological standard. For beekeepers, the key message is therefore a word of caution: it is better to strengthen the foundation of the apiary and the quality of resources than to rely on additives that have not yet been sufficiently validated.
There is a simple and fairly reliable method for adjusting (calibrating) the refractometer: using extra virgin olive oil.
According to a test conducted by Dr. Werner von der Ohe from the Bee Institute in Celle, calibration of the refractometer with extra virgin olive oil at a temperature of 20 ºC is possible. Tests using three different extra virgin olive oils showed a maximum deviation of 0.08 % (see the article published below in ADIZ – die biene – Imkerfreund).
Diseases and pests pose a major threat to the health of colonies and require early detection and appropriate management. This overview presents the main diseases and pests affecting honey bees. The following information is based on official fact sheet 2 from the Swiss Bee Health Service.
It sticks, it stains, and we sometimes curse it when inspecting our beehives; yet it gives beekeeping one of the fragrances that contribute to its charm and is also an increasingly sought-after product, representing a growing source of income for beekeepers. Although omnipresent in our hives, propolis is the result of a harvest and a processing effort that is anything but simple for the bee. A demanding but indispensable task; for this resinous substance, whose medicinal properties have been appreciated since Antiquity, fulfils multiple functions within the colony.

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© africa-studio.com (Olga Yastremska and Leonid Yastremskiy)
When a colony appears to be lacking in vitality, it can be tempting to turn to a protein paste or a homemade recipe. However, natural pollen is more than just a source of protein, and scientific studies do not support the routine use of these supplements. In Switzerland, Agroscope summarizes this caution as follows: protein-based supplemental feed for bees rarely offers benefits, but often has drawbacks.
The rhododendron is an emblematic melliferous plant and ranks among the most important nectar sources in mountain regions. In Switzerland, two species are present: Rhododendron ferrugineum, which grows on acidic soils, and Rhododendron hirsutum, which prefers calcareous soils.
Heathers are small, bushy shrubs belonging to the Ericaceae family that grow on nutrient-poor soils. There are many species – several hundred – whose flowering periods extend almost throughout the entire year: June to August for Erica tetralix, June to October for Erica cinerea, June to December for Erica arborea, August to October for Calluna vulgaris, September to November for Erica multifolia, November to April or February to July for Erica carnea, etc.
The articles proposed by experienced beekeepers for the training of younger colleagues remind us that colonies should be carefully insulated for overwintering.
However, a recent German study reports that there is no advantage, in terms of brood initiation or colony development in spring, associated with sophisticated insulation of the colony. The dogma of insulating the winter cluster is thus being challenged. By discovering the precision of the winter cluster’s thermoregulation, the inquisitive beekeeper is prompted to reflect on their beekeeping practices.
Most bee races do not leave the hive if the outside temperature is below 10 to 12 °C.
Hives should not be opened if the temperature is below 15 °C. If an inspection lasts too long, there is a risk of chilling the brood, which can lead to its death or to diseases. Calm movements are essential, and one should never lose control of the situation!
The economic benefit for beekeeping operations of removing drone brood as a complementary measure
to control varroa
Since the first article, we have received a large number of requests for clarification regarding the practical implementation
in the hive, followed by many very positive comments on the effects of this “mechanical treatment,” which poses no risk to bees or the queen. After recalling the context of the study and the way it was conducted, we now present the complete results over a two-year period.

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Reducing the space available to a colony is often presented as a way to keep bees “warm”. In reality, the issue is more nuanced: bees do not heat the whole hive, but organise a functional nest around brood, the winter cluster and food stores. This article explains when reducing space can support the colony — and when it risks replacing observation with an overly rigid rule.

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By 2024, Switzerland had already warmed by about 2.9 °C relative to pre-industrial conditions (1871–1900) — more than twice the global average. That is roughly the warming level the CH2025 climate scenarios describe for Switzerland in a global 1.5-degree world. For beekeeping, adaptation is therefore not a task for the future. However, the key issue is not simply that everything is happening earlier. Flowering, nectar flow, colony development, Varroa reproduction, and suitable treatment windows are shifting at different rates and no longer coincide as reliably. Earlier flowering does not guarantee a nectar flow; strong colony development simultaneously creates more space for Varroa to reproduce; and a warm fall delays the brood-free period. As a result, the beekeeping calendar does not lose its value, but it does lose its predictive power. This article combines the official Swiss climate scenarios with findings from plant biology, ecology, and bee research, consistently distinguishing between established findings, plausible inferences, and open questions.

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Winter bees, often referred to as diutinus bees in the scientific literature, are long-lived workers adapted to ensuring the colony’s survival during winter. They do not constitute a distinct caste, but rather a seasonal form of the worker bee, which develops as the colony gradually shifts from a growth phase to a conservation phase. Understanding this transition provides a better understanding of what happens in the apiary between the end of summer, winter, and the spring revival.
Bees that build their own combs are less inclined to swarm, as the pheromones released during wax production inform the queen about the increase in the volume of the hive.
The wax originates from specialized glands located in the abdomen of young wax-producing worker bees approximately 12 days old, developing in an environment with temperatures between 33 °C and 36 °C. These workers consume large amounts of sugar for wax production, about 7.5 kg to produce 1 kg of wax. The wax appears in the form of small scales weighing less than 1 mg. Using her hind legs, the bee brings them to her mandibles, kneads them, and assembles them
Under the screened bottoms of our hive floors, it is advisable to place a drawer. Various debris, poorly stored in the cells, end up on this board and can sometimes provide information without having to open the hive.
Although the wax moth has a very bad reputation, beekeeping manuals often underestimate the damage caused by the larvae of the greater wax moth Galleria mellonella and the lesser wax moth Achroea grisella, which consume everything that comes within reach of their mandibles (wax, food reserves, pollen stores, wood, polystyrene—not to mention the brood!). It is commonly assumed that strong colonies can defend themselves very easily against wax moths … but this is not always the case. In fact, wax moths reproduce very rapidly, and the larvae of the greater wax moth feed on capped brood, sheltered from counter-attacks by worker bees. As a result, the bee population may decline, as may the colony’s defences against this formidable pest. Selection for overly gentle bees and the pheromonal mimicry of the wax moth, which imitates queen pheromones, can ultimately lead to colony collapse.
Despite these harmful effects, it should be emphasised that wax moths fulfil an important hygienic function, especially in the context of “poor” beekeeping practices. They destroy abandoned bee nests and return the recovered materials to the natural cycle. In this way, pathogens responsible for diseases such as American foulbrood and European foulbrood are also eliminated.
Do you recognize the diseases?
Take the test: try to assess the diseases!
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A colony may replace its queen for several reasons: advanced age, insufficient egg-laying, poor fertilization, injury, or general weakness. But a recent study suggests that another factor may sometimes be at play: a severe viral infection in the queen could impair her reproductive function and alter her pheromonal signal, to the point of triggering supersedure.
The insulation of beehives during winter has long been the subject of debate within the beekeeping community. While some studies emphasize its beneficial effects on reducing energy consumption and improving colony survival (St. Clair et al., 2022; Alburaki & Corona, 2021), others highlight its limitations and potential side effects, particularly when it disrupts the natural thermoregulation mechanisms of the superorganism (Mitchell, 2023; Minaud et al., 2024).
The purchase and sale of bee colonies must be carried out transparently, with knowledge of their health status, origin, and treatment history. The summary below is based exclusively on the official checklist 4.11 of the Swiss Beekeeping Service (BGD/SSA) and refers to the complete documents for further information.
Proper storage of frames is an essential part of beekeeping hygiene and helps prevent damage caused by pests, mold, or contamination. The checklist describes appropriate storage methods and precautions to be taken. The following information is based on the official checklist 4.4.2 of the Swiss Bee Health Service.
Hyperthermia is a biotechnical method for reducing Varroa infestation in which brood is deliberately heated to kill mites. It exploits the differing temperature sensitivity of honey bee brood and Varroa mites. The following notes are based on the official Practical Guide 1.6.3 of the Swiss Bee Health Service.
Beekeepers use many methods to unite bee colonies, and some very violent ones demonstrate a clear lack of respect for the bees. The only advantage of such methods is their speed, but they pay little attention to what happens to the queens, the foragers, and the bees in general. The gentlest and most commonly used method is very probably the “superposition” method. This procedure is relatively simple, accessible to both beginner and experienced beekeepers, and if the few rules discussed in this article are respected, the uniting of colonies should proceed without any problems.
The Geneva-based naturalist became one of the greatest specialists on bees of his time without being able to see them: he was blind and conducted his research through the eyes and hands of his faithful assistant, François Burnens.
Varroa treatment in mating nuclei places specific demands, as these units usually consist of small bee populations with a particular function. An adapted approach is required to ensure treatment effectiveness while avoiding risks to queens and brood. The following notes are based on the official Practical Guide 1.6.5 of the Swiss Bee Health Service.
Propolis is a complex material, composed mainly of resins derived from various plant species, but also containing variable amounts of wax produced by the bees themselves.
Good apicultural practice: The advantage of spring nucleus colonies over summer nuclei is that they allow a first assessment of queen quality prior to overwintering, owing to an additional month of egg-laying activity.
Im չափfect clones without sons, males without fathers but with a maternal grandfather, twins through their fathers, sisters through their mothers, daughters as full or half-sisters …
Let us get straight to the point: honey bee genetics is truly unusual! With bees, one must set aside what is known from human reproduction, where each parent contributes half of the chromosomes—the mother via the egg, the father via the sperm. This is not the case in bees!
The question comes up every year: should colonies be fed at the end of winter or not? The answer needs to be nuanced. Food reserves have a direct impact on the queen’s egg-laying activity. It is well known that generous reserves and regular nectar inputs clearly stimulate egg laying, and when pollen is abundant, nurse bees operate at full capacity.
Measuring the natural mite fall is a simple method for assessing varroa infestation without treatment intervention. It provides orientation on the infestation level and supports decision-making regarding further measures. The following information is based on the official Practical Guide 1.5.1 of the Swiss Bee Health Service.
Beeswax is a biologically relevant compartment within the colony. It can act as a reservoir for pathogens and chemical contaminants and influence brood development and certain colony performance parameters. Structured management of the wax cycle, including regular renewal of brood combs, therefore appears to be a scientifically based lever for reducing the cumulative accumulation of risks.

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The Asian hornet is spreading gradually in Switzerland and is becoming an increasing concern for beekeepers. It is therefore essential to recognize it reliably, distinguish its nests, and respond appropriately when in doubt. This article provides practical guidance on how to identify the species, report a suspected case correctly, and understand which protective measures at the apiary are currently the most realistic.
Among Asian knotweeds, the Japanese knotweed, whose scientific names are Fallopia japonica, Polygonum japonicum or Reynoutria japonica, is the most widespread. It belongs to the Polygonaceae family and originates from the Far East (eastern China, Japan, Korea). Introduced to Europe in the 19th century for its ornamental and forage properties, it proved to be invasive, spreading rapidly across large areas, particularly in moist environments.

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In June, the apiary remains highly active, but the logic of the season is already beginning to shift. Between nectar flows, lack of space, swarming, colony splitting, transhumance, and Varroa monitoring, the beekeeper must both manage still very strong colonies and prepare for the rest of the summer. This article outlines the main issues and tasks for the month of June in a way that is suited to the realities of beekeeping in Switzerland.

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Honey is far more than a concentrated sugar solution: it is the result of a complex biological process shaped by botanical origin, processing by the bees, and the conditions of ripening and storage. This reference article explores its composition, physical and microbiological properties, quality, authenticity, health risks, and uses, while distinguishing well-established evidence from remaining uncertainties and common overstatements.
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RNA interference offers a new approach to varroa control: instead of killing the mite directly, it aims to block the expression of an essential parasite gene. This article summarizes what is currently known about vadescana/Norroa, early laboratory and field trials, and the remaining questions for honey bees, non-target organisms, and beekeeping in Switzerland.
Berberis vulgaris is the scientific name of the barberry, a shrub that owes its name to a lightly fermented beverage made from its red berries, which resembles wine. However, its interest for beekeepers lies mainly in its flowers, which produce very abundant nectar during the months of May and June.
This study entitled “Safeguarding pollinators and their values to human well-being” estimates that “75%
of major crops – cereals, fruits – depend on pollination by animal species”. Their disappearance would lead to a loss of 5 to 8% of global agricultural production, which would trigger an increase in potential resurgences of diseases linked to our dietary balance (heart disease) and would increase “by 1.4 million per year, the number of additional deaths”.
May disease is not an infectious disease, but a metabolic disorder that mainly occurs in spring when the brood is developing rapidly. It is linked to a lack of water in the colony. The following information is based on official memo 2.9 from the Swiss Bee Health Service.
Left or right? Like humans, bees also show a preference. Australian researchers have recently discovered that bees can have a left- or right-side preference—affecting their flight decisions in order to avoid obstacles.
The human brain consists of two hemispheres that are roughly symmetrical in terms of morphology. Functionally, however, there is marked asymmetry: most of us are right-handed, and everyday objects are designed for right-handed users—the pair of scissors, the tape measure, the computer mouse, the guitar, the graduated jug, the fold-down writing tablet attached to an auditorium chair, the stock of a hunting rifle, the boomerang… not to mention the convention of the handshake.
A recent study reveals that while 50% of bees are not lateralised, 25% are right-biased and 25% are left-biased. One hypothesis proposed to explain this phenomenon is that swarm flight strategies are optimised, thereby avoiding a proportion of collisions.
The implementation of a sentinel bee project, through its epidemiological dimension, constitutes the only approach capable of shedding light on the causes of the significant losses that have been affecting bee populations for nearly twenty years. Moreover, such a project makes it possible to assess the state of the environment in which colonies live—an environment that is also our own—in which the bee, owing to its sensitivity to toxic contaminants, plays an early warning role.
Principles, method and key control points
Inspecting a hive is neither a trivial action nor an automatic routine. Each opening disrupts the colony’s internal balance: brood temperature, humidity, spatial organisation and chemical communication. Field observations show that after an inspection, a colony may need 24 to 48 hours to fully restore its functional equilibrium.
The modern beekeeper therefore does not inspect out of curiosity, but to address a clearly defined objective. A sound inspection is based on a simple logic: observe → diagnose → decide → close, aiming to obtain the maximum amount of information with the minimum level of intrusion.
Collecting pollen is no small task : bees therefore use several senses and different techniques to choose it.
A study involving Inra, Acta and the ITSAP-Institut de l’abeille examines the physiological mechanisms involved in the winter survival of honey bees. The researchers showed that a protein with antioxidant properties, vitellogenin, is associated with a 30% increase in the probability of colony survival during winter.

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How does a honey bee colony keep its brood within a narrow temperature range, cope with cold and heat, and how much does the hive itself matter? This reference article explains the biological and physical mechanisms of thermoregulation, critically examines insulation, humidity, ventilation and insulated partitions, and distinguishes well-supported practical recommendations from questions that science cannot yet answer with certainty.
Those who have already used the excuse that “housekeeping is not in my DNA” to try to avoid this chore may not have been entirely wrong. Indeed, at least in honey bees, certain shortcomings in hive hygiene appear to be linked to the overexpression of genes that impairs the detection of odors released by diseased or dead larvae.
Unmarked queens can sometimes be very difficult to locate. We recommend having marking equipment available during every colony inspection, so that unmarked queens can always be marked when encountered. This requires a good eye, and not all beekeepers necessarily have the ability to reliably spot queens. When a nucleus colony needs to be formed or a queen introduced, the beekeeper is then often faced with a difficulty.
The Asian hornet has continued its expansion since its arrival in France 20 years ago.
In 2024, it has reached the borders of Valais, and the Saint-Maurice gorge will most likely not constitute an insurmountable barrier for this highly well-adapted invader.
Propolis is a natural substance produced by bees and plays an essential role in the hygiene and stability of the colony. The fact sheet describes its origin, functions, and importance in the colony. The following information is based on the official fact sheet 4.1.1 from the Swiss Bee Health Service.
The division of a colony is carried out for various reasons :
There are many different and very simple methods for dividing hives.
The method described below works well :
During foraging, honey bees exhibit remarkable cognitive abilities. However, pesticides and heavy metals disrupt neuronal communication, impair foraging behaviour, and ultimately place the entire colony at risk.
The effects of malnutrition in bees are well documented. There is an interaction between individual bees and the colony, and individual problems are reflected in the adult population and brood, which are reduced both qualitatively and quantitatively. Antonio Gómez Pajuelo is recognised as one of the world’s experts on bee nutrition. He conducted a survey involving 166 beekeepers.
by DALILA BOVET
Personality in animals can be discussed when consistent individual differences are observed over time and expressed across different contexts. Some bees prove to be more attracted to novelty than others. These behavioural differences are based on genetic variations. Can we therefore conclude that bees have a personality?
Swarming is a natural reproductive behaviour of the honey bee colony, but in beekeeping it often leads to losses, extra work and uncertainty. This article explains why swarming tendency should not be interpreted too quickly as a genetic problem of the queen, but as the result of interactions between genetics, queen age, nectar flow, available space and colony management.
Summer treatment is a central component of varroa control in Switzerland. The Liebig dispenser is a widely used method for the application of formic acid (Formivar) and requires correct handling to ensure both efficacy and tolerability. The summary below is based exclusively on the official practical guide (BGD/SSA) and refers to the complete instructions.
Deciduous tree or occasionally evergreen, monoecious and wind-pollinated, with inconspicuous, greenish and unisexual flowers. The male flowers are pendulous and grouped in catkins, whereas the female flowers are rounded and hardly visible, developing into acorns (nut fruits).
The Asian hornet is an invasive species that poses a significant threat to honeybees and other insects. Its presence requires increased vigilance due to its ability to spread rapidly and its impact on colonies. The following information is based on the official checklist 2.7 of the Swiss Bee Health Service.
The tree of heaven (Ailanthus altissima), also known as the Japanese varnish tree, is a tree belonging to the family Simaroubaceae that can reach a height of 25 to 30 metres. It originates from East Asia and was deliberately introduced into Europe in the 18th century for ornamental purposes, but above all in connection with silk production, as the plant served as a food source for a moth, the ailanthus silkmoth (Samia cynthia), which was intended to replace the mulberry silkworm (Bombyx mori).
In the honey bee, as in all insects, the haemolymph fills the entire internal cavity protected by the cuticle. This cavity is called the haemocoel. There is no network of veins and arteries: the organs are bathed in the haemolymph, which supplies the organism with the necessary components. It is referred to as an extracellular fluid. A long tubular vessel, the heart, pumps the haemolymph and ensures the circulation of the fluid throughout the organism.
The creation, as early as spring, of several nucleus colonies from a small six-frame colony is highly profitable and straightforward, provided that the beekeeper monitors the development of the parent colony, transfers the brood frames onto two stacked boxes, and that frequent syrup feeding stimulates a particularly prolific queen.
A very recent American study challenges the dogma of queen rearing that has existed since the late 1890s.
From their initiation into queen rearing, beekeepers have been taught that the factor determining the caste of larvae originating from fertilized eggs is the duration of feeding with royal jelly.
If a larva is fed exclusively with royal jelly during the six days preceding cell capping, the adult insect that emerges on day 16 will be a (virgin) queen. By contrast, if the same larva is fed with a mixture of honey and pollen during the last three days before capping, a sterile worker will emerge on day 21. A very recent American study published in 2020 by the Royal Society challenges this long-established dogma.

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Blessed is the one who manages to escape the bee that is chasing him. The flight of the honey bee is a mechanical and sensory feat: flexible wings, an elastic thorax, specialised muscles and rapid vision work together. This article summarises what is known about this flight mechanics and shows what beekeepers can take from it to better interpret activity at the hive entrance, bee orientation and certain warning signs in the apiary.
What is the approximate brood volume as a function of the queen’s egg-laying capacity? How many brood frames does the queen need in order to lay eggs without space limitation?
The number of plant species producing nectar and pollen is very large; in Switzerland, several thousand are recorded. However, they do not all have the same beekeeping value, and only a few hundred are actually visited by bees. Among these, barely around thirty have a truly high beekeeping value, about seventy are considered secondary, and the others are of lesser interest because they are less common or occur only sporadically.
The structure used to house bees has evolved considerably over time. Early beekeepers used pottery vessels, hollow walls, sections of tree trunks, or woven baskets, sometimes covered with dried earth. A few examples of these systems are still in use, but the framed hive system has gradually become the standard.
Emergency treatment in hive bodies refers to exceptional measures that may be applied in the event of an acute threat to a honey bee colony. It is not intended for routine colony management, but for short-term intervention in critical situations. The following notes are based on the official Practical Guide 1.7.1 of the Swiss Bee Health Service.
A bee colony requires tranquillity throughout the beekeeping season, and probably even more so during the winter months. If an overly curious beekeeper opens the hive indiscriminately, the constantly disturbed colony becomes stressed. The behaviour of this superorganism changes: energy consumption increases, the reserves of its valuable fat body are depleted, immune defences against varroa and the viruses it transmits are inhibited, the development of the worker population and brood slows down, and the colony ultimately collapses into a vicious downward spiral (► Infernal cascade: Chronicle of a foretold death).

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The naturally resistant bee populations observed on Gotland, and later in other regions of Europe and elsewhere, have had a profound impact on thinking about the Varroa mite. They demonstrate that sustainable survival without treatment can emerge through natural selection, without being reduced to a single mechanism or a universally applicable solution. The insights they offer—as well as the limitations they impose on interpretation—must therefore be approached with the same rigor.
Unmarked queens are sometimes very difficult to locate. We recommend having marking equipment available during every colony inspection so that unmarked queens can always be marked. This requires a good eye, and not all beekeepers necessarily have the ability to spot queens. When a nucleus colony has to be formed or a queen introduced, the beekeeper is then often faced with difficulties.
by FRANÇOIS SAVATIER
Traces of beeswax found on pottery sherds suggest that beekeeping was already mastered at the beginning of the Neolithic period.
Acquired and controlled through evolution, a degree of “disorder” in the nanometric structure of the petals of many flowers allows for more efficient pollination.
Researchers have discovered that some flowers have developed an additional strategy to encourage insects to forage on them. An ultraviolet halo effectively attracts pollinators.
Image: Ursinia speciosa, like other flower species, produces a blue halo on its petals to attract pollinators. © Edwige Moyroud
In November, the apiary gradually enters its winter dormancy, but not all colonies are necessarily brood-free yet. The key is to observe without disturbing the bees, secure the hives, and check the food stores. The decision to treat for varroa mites in winter is not based on a fixed date, but rather when the absence of capped brood is confirmed.
Robbing is a complex behaviour that involves more than the visible loss of a colony’s food stores. It should not be underestimated, as its consequences can be serious for the affected colony and for the health of the apiary as a whole. This article explains how robbing develops, how foreign bees are recognised, and which measures can help prevent or stop it.
Pollen is far more than a simple source of protein: its composition, digestibility and nutritional value vary considerably from plant to plant. From the pollen pellet as collected to the bee bread stored in the cells, this article examines what science establishes about its essential role in the development, health and survival of colonies, and draws out the implications for beekeeping practice. Finally, it distinguishes this vital importance for bees from the benefits attributed to pollen in humans, assessing its nutritional value, the health claims made for it, the limits of the clinical evidence and the risks associated with its consumption.
The first difficulty for a beginner beekeeper is choosing a hive. There are many systems, and all of them have their advantages and disadvantages, their supporters and their critics. It is important to choose hives carefully so that the colony develops well, production is satisfactory, overwintering proceeds properly, and beekeeping remains a pleasure. Dadant, Langstroth, Voirnot, Alsacienne, WBC, Warré, Layens, Simplex, Zander, National … and all the others: which is the best hive? How can one make a decision when starting out in beekeeping and the opinions of the “experienced beekeepers” diverge?
Adherence to recommended treatment regimens against Varroa destructor improves the winter survival of honey bee colonies. Key findings:
Young bees aged 12–19 days (wax-producing bees)* produce small wax scales using their wax glands, which are located in the last four abdominal segments. The process of wax production is complex.
February is a pivotal month for the colony. Day length increases, and the queen gradually resumes laying. This resumption remains fragile: stores can become critical, temperatures are unstable, and any prolonged opening can chill the brood. The beekeeper acts mainly through observation, anticipation, and preparation, keeping interventions to a minimum.
Last year, the publication of specialised articles on the contamination and adulteration of beeswax shook the beekeeping community. Among beekeepers, awareness has grown regarding the importance of the condition of wax: indeed, both the quality of our products and, of course, the health of our colonies depend on it (1).
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Bees do not predict the end of the world, but they can reveal important signals about the state of the environment. This Belgian report shows why honey bees can act as sentinels for certain environmental contaminants, while wild bees are more informative about habitat quality. The key is not to draw quick conclusions from a single apiary, but to build collective, rigorous and well-documented monitoring.
Canadian fleabane (Conyza canadensis or Erigeron canadensis) and annual fleabane (Erigeron annuus or Stenactis annua) are annual herbaceous plants belonging to the Asteraceae family and originating from North America. They exhibit all the characteristics of invasive nectar-producing plants: exotic origin; rapid growth; production of a large number of wind-dispersed seeds that can be transported over several kilometres; ability to germinate in the smallest crevices; and strong attractiveness to pollinating insects, including honey bees.

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The drone is the male of the honeybee. Its caste depends on the type of egg, the larval diet, and the type of cell. It develops from an unfertilized egg laid in a drone cell. It is the largest insect in the colony, recognizable by its stocky body, hairy thorax, very large eyes, and noisy flight. Its primary role is to transmit its mother’s genetic material during fertilization.
Which medications and treatments are approved in Switzerland for varroa mites, and what key points should be kept in mind before treatment? This summary outlines the main approved treatments and highlights a crucial point for beekeepers: when using medication, the supers intended for honey harvest must be removed, and the product’s official instructions must always be followed.
During May, a simple way to increase one’s apiary or to prevent swarming in an overly strong colony is to create a package bee swarm.
For a long time, scientists believed that an organism’s genetic code alone determined its biological characteristics, its development, and its adaptation to its environment. Under the influence of various chemical or “external” factors, epigenetics can not only modify an organism’s appearance or behaviour, but can also transmit some of these modifications to subsequent generations…
Saffron, a highly valued culinary spice, consists of the dried stigmas of a crocus species, Crocus sativus, also known as cultivated saffron. It is a perennial herbaceous bulb plant whose flower has six tepals (tepals = floral organs similar to petals), which are mauve with violet streaks.
Bee packages – all similar, yet very different in their composition and use.
The preparation of bee packages is not a common practice in beekeeping, as it requires specific expertise that is rarely taught in beekeeping schools. Many online videos fail to provide all the information surrounding this technique, which often leads to unsuccessful outcomes when beekeepers attempt to put it into practice.
Replacing old frames is part of the classical recommendations in beekeeping. In practice, however, this measure is often postponed, mainly for economic reasons or due to underestimation of its biological effects. Yet several experimental studies show that comb age directly influences bee morphology, colony demographic dynamics, honey productivity, as well as the accumulation of contaminants in the hive (Berry & Delaplane, 2001 ; Abd Al-Fattah et al., 2021 ; Taha et al., 2021 ; Meng et al., 2025).
Insects are not insensitive automatons: bees, wasps, flies, or ants display remarkable cognitive and emotional capacities under laboratory conditions.
Recent experiments suggest that bumblebees may experience optimism, joy, and possibly even pain. These findings raise important ethical questions concerning the treatment of insects in laboratory settings, their breeding, and the use of pesticides.
Based on the videoconference by Prof. Joseph Hemmerlé on 11.01.2025, School of Agriculture Châteauneuf / Sion
Pollen is an essential resource for bees, but it is also of significant interest to humans. This presentation highlights the roles, characteristics, and applications of this fascinating biological material.
Queen multiplication using the Laurenz method is a breeding technique that does not require grafting. This guide outlines the conditions, procedure, and limitations of this method. The following information is based on the official guide 4.6.2 published by the Swiss Bee Health Service.
Total brood removal is a biotechnical measure for reducing Varroa infestation without the direct use of varroacides. The procedure deliberately intervenes in the brood cycle and requires stable, high-performing colonies. The following notes are based exclusively on the official Practical Guide 1.6.4 of the Swiss Bee Health Service.
When one speaks of alfalfa, one first thinks of cultivated alfalfa, Medicago sativa, a perennial herbaceous plant native to the Middle East with blue-violet flowers; however, there are other species with flowers in different shades, ranging from yellow to white, via green and brown. All are highly melliferous.
Too often, the drawer is used solely to detect the presence, more or less abundant, of natural debris from dead varroa mites. Yet the drawer is a mirror of the life of the colony just above it… If the beekeeper takes the time to examine it regularly, the observed elements, waste, fragments, and other residues provide valuable information about colony dynamics and health. Examination of the drawer must always be correlated with the beekeeping calendar: the interpretation of a drawer inspected in summer will be very different from that of the same drawer opened at Christmas.
From the honey super still on the hive to the sealed jar, every step of the harvest determines the quality of the final product. This article follows the complete process : assessing ripeness, extracting under the right conditions, managing crystallisation and filling correctly. Mistakes made early in the process cannot be corrected later — which is why every decision is explained together with its practical consequences. An article for beekeepers who want to understand why, not just how.
Based on the videoconference by Prof. Joseph Hemmerlé on 11.01.2025, School of Agriculture Châteauneuf / Sion
Swarming is a natural phenomenon at the core of honey bee colony dynamics. Through this process, part of the colony, led by the old queen, leaves the hive to establish a new nest. For beekeepers, swarming represents a challenge, but it also constitutes an opportunity for colony renewal. Thanks to the in-depth research and careful observations conducted by Professor Joseph Hemmerlé, it is possible to gain a better understanding of the biological, ethological, and environmental mechanisms underlying this fascinating behaviour.
Ask your question about beekeeping. The more detailed your description, the more helpful and relevant the answer will be to your specific situation at the apiary.
Controlled melting of frames (e.g., to recover wax or treat used frames) produces clean wax and facilitates the removal of undesirable substances. This summary is based exclusively on the official guidance document 4.4.1 (BGD/SSA).
The cells in the comb are not only used for storing honey or raising brood. A study shows that bees also use them as a place to rest or even sleep. By observing the movements of the abdomen, it is possible to distinguish between a sleeping bee and a bee busy warming the brood, which opens up new perspectives on the organization of life in the colony.
Die Grundregel ist einfach: Oxalsäure wirkt vor allem außerhalb der Brut, während Ameisensäure dann sinnvoll ist, wenn noch verdeckelte Brut vorhanden ist. Im Herbst weichen bestimmte Situationen jedoch von der üblichen Logik ab: Bevor man über das Mittel nachdenkt, muss man sich zunächst fragen, ob der Bienenstock noch winterfähig ist und welche Maßnahme zu diesem Zeitpunkt der Saison noch sinnvoll ist.
The introduction of queens always entails a significant risk for the introduced queen. Some publications report failure rates of up to 50 %. For successful acceptance, all conditions must be optimal. The highest success rates are achieved when a young colony with a young queen is combined with an older colony. In general, favourable conditions must be deliberately created in order for a colony to accept an introduced queen.
Would it not be interesting to preserve the alleles of the most productive or most important queens after their death? It is therefore not surprising that the cryopreservation of drone semen has been the subject of research since the 1970s. The risk of losing natural biodiversity is another motivation for this research.
Queen multiplication using the Mini-Plus system is a recognized method for breeding, fertilizing, and preserving young queens. It allows for precise monitoring of queens in small colonies. The information below is based on the official checklist 4.6.1 from the Swiss Bee Health Service.
The location of the apiary is undoubtedly the most important choice. It will determine, of course, the harvest, the amount of time you will spend with your bees, and the various savings you will be able to make.
The winter survival of bee colonies is a topic that divides beekeepers because of contradictions, preconceptions, hypotheses and differing viewpoints. The experienced beekeeper relies on experience and closely monitors weather variations from November to March in order to estimate the timing of the resumption of egg laying by the queen and the volume of brood that must be kept warm. The beginner beekeeper, by contrast, is confronted with the questions raised by worried colleagues: should colonies be fed during winter or not?
This article provides a structured introduction to queen rearing for beekeepers who want to become self-sufficient in their apiary. It presents, in an educational manner, the necessary equipment, rigorous planning of the rearing schedule, and a simple, proven method suitable for an annual series. The aim is to enable everyone to produce their own high-quality F1 queens, while drawing on the selection work carried out by the Breeding Instructors.
The evaluation or selection of colonies does not concern breeders alone; it is also important for every beekeeper from the perspective of bee health. As a preventive measure, the ultimate aim is to retain only healthy and strong colonies within an apiary. This selection applies both to productive colonies and to young colonies.
This article begins by examining the life cycle of the varroa mite to identify the two windows of opportunity available to a colony in dealing with the parasite, and then reviews the known mechanisms and methods for evaluating them. It constitutes the practical and methodological component of the special report on varroa resistance.
The FAM dispenser is an approved device for the treatment of honey bee colonies with formic acid and is primarily used for varroa control. Correct application is essential, as formic acid is highly corrosive and improper handling poses risks to both humans and bees. The following information is based on the official Practical Guide 1.2.3 of the Swiss Bee Health Service.
The parasitic mite is decimating beehives. Biologist Paul Page has shown that an Asian forager has found a countermeasure through a system of “altruistic suicide.” A model that its European cousin could follow?
What beekeepers should know about exposure, immune reactions, and protection
Among beekeepers, bee stings constitute a frequent occupational exposure, often trivialized and traditionally associated with the idea of tolerance acquired through repeated exposure. This widely held representation is based on genuine empirical observations, but it only partially reflects the documented clinical and immunological reality.
August marks a turning point at the apiary: the harvest is coming to an end, while decisions regarding varroa mites, food stores, and colony selection are already setting the stage for winter. This article summarizes the month’s practical priorities, common mistakes to avoid, and key considerations that determine the quality of the winter bees.
The rowan is a deciduous tree that is very widespread in Valais, where it grows naturally in woodlands but is also widely planted in parks and gardens, notably to form living hedges that are highly valued for their foliage and decorative fruits. Several species of rowan exist; the most common are the rowanberry or mountain ash (Sorbus aucuparia), the service tree, also known as sorb apple tree (Sorbus domestica), and the whitebeam (Sorbus aria).
Pheromones are key elements of animal communication: they are released to convey specific messages such as sexual attraction, aggression, recognition of conspecifics, etc., to members of the same species.
Figure: Proboscis extension: appetitive response of an immobilized bee to a sugar-solution reward that has contacted its antennae. Bees exposed to pheromones of different significance modify their proboscis extension behavior, thereby demonstrating the impact of these pheromones on the evaluation of the received food reward. © Martin Giurfa
Castanea sativa Miller is the scientific name of the chestnut tree. It is a majestic, long-lived tree that can reach up to 40 metres in height and 15 metres in circumference. As it does not tolerate calcium and its salts, in particular carbonates (limestone), it is found on acidic soils; this explains why it is not evenly distributed in Valais.
There is no single correct beekeeping practice. However, there are rules based on common sense and on the experience of seasoned or professional beekeepers. While the queen is indeed the driving force of the colony, hive cleanliness, control of varroa mites, swarm-prevention strategies, the location of the apiary, and winter preparation are equally important for the healthy development of the colony with a view to achieving a good honey harvest.
The parasite capable of chemically mimicking two bee species
Researchers from the Institute for Research on Insect Biology (CNRS/Université François Rabelais de Tours) and the “Bees and Environment” laboratory of Inra, in collaboration with American and Chinese colleagues(1), have shown that Varroa destructor, a parasitic mite of bees that is able to imitate the chemical composition of the cuticle(2) of its host, can also modify this composition depending on the species it parasitises. This remarkable adaptive capacity could explain how this parasite of the Asian honey bee was able to colonise the European honey bee during the 20th century, thereby contributing to the decline of the species. These findings were published on 3 June 2015 in the journal Biology Letters.
Invasive plants are non-native plant species that have been introduced intentionally or accidentally by humans outside their area of origin and that reproduce and spread extensively, to the detriment of native species. Most often originating from other continents, they profoundly disrupt ecosystems and reduce local biological diversity.
Ideally, high-quality honey has a water content not exceeding 17.5%. This allows the beekeeper to sell honey of high quality that does not ferment once it reaches the consumer.
Today, many so-called “probiotic” products are available on the market, intended to improve our health or sometimes even that of our livestock, such as bees1. Probiotics are formulations based on microorganisms that maintain beneficial relationships with their host. The purpose of this article is to provide elements for reflection in order to understand how such products might potentially contribute to combating diseases of the honey bee.
The eggs laid by the queen pass through several developmental stages. The first, which lasts barely three days, is the most critical, as it corresponds to the development of the nervous and digestive systems of the larva inside the egg. After hatching, the larval stage begins with the emergence of a larva without antennae, eyes, legs, or wings, which is fed exclusively on a diet of royal jelly for three days.
(Photo: kleinewelt)
Biotechnical control methods are increasingly being promoted in the fight against varroosis. Among these, techniques that induce significant interruptions in brood development are often discussed. However, such methods are still rarely used by beekeepers, except on an experimental basis. This article analyses the relevance of brood interruption and the different techniques derived from it.
Dynamic colony management is a pragmatic approach to colony maintenance, adapted to their health status, available resources, and rate of development. It combines brood extraction, nucleus creation, swarm prevention, queen selection, and resource optimization according to seasonal needs. This summary is based on the official checklist 4.12 of the Swiss Beekeeping Service (BGD/SSA).
Die Nestsuche durch Triangulation ist eine gezielte Methode zur Lokalisierung von Nestern der Asiatischen Hornisse. Sie basiert auf systematischen Beobachtungen der Flugrichtungen einzelner Tiere und unterstützt die Behörden bei der Bekämpfung dieser invasiven Art. Die folgenden Hinweise basieren auf dem offiziellen Merkblatt 2.7.2 des Schweizerischen Bienengesundheitsdienstes.
Although substantial research has been conducted on the causes of colony collapse disorder in the European honey bee Apis mellifera, there has been increasing evidence over the past two decades that another pandemic affecting both domesticated and native bees is emerging. This pandemic is the result of the spread of fungal pathogens of the genus Nosema.
Emergency varroa treatment in Swiss hives is a measure intended for colonies that are too heavily infested, in order to sanitize them within one day. Official Practical Guide 1.7.2 of the SSA / BGD describes a technique corresponding to the artificial swarm with queen method, with rehousing of the colony on frames with foundation. The document links the intervention to the natural mite drop, to treatment on a colony without capped brood, and to the complete renewal of all frames.
Dribble treatment with oxalic acid is an established method in Switzerland for winter varroa control. Broodless colonies (or colonies with only open brood), correct preparation of the solution, and strict protective measures are essential, as oxalic acid is strongly irritating and harmful to health. The summary below is based exclusively on the official practical guide (BGD/SSA) and refers to the complete instructions/package leaflet.
Robbing is undesirable behavior in which bees from other colonies enter a hive to steal its reserves. It often occurs during periods of food shortage or following beekeeping activities. The following information is based on official guideline 4.8.3 from the Swiss Bee Health Service.
Members of an “Apis mellifera” colony share the same olfactory signature, emitted by their cuticular hydrocarbons and linked to a specific intestinal bacterial flora.
Like any living organism, the bee is prey to various parasites and pathogenic germs: every colony contains a broad and diverse range of microbes. Yet not every colony becomes ill as a result! This is because, like all living beings, bees have developed various means over the course of evolution to resist them: this is what is known as immunity. Bees, like humans, possess an immune defence system, but with some differences.
The glandular system, the circulatory, respiratory and nervous systems, as well as the digestive, excretory and reproductive systems, will be presented in turn in the following fact sheets. When addressing the internal anatomy of the honey bee, an anatomical plate is more informative than lengthy explanations.
Our bee populations are not able to defend themselves against varroa on their own. They therefore rely on you, beekeepers, to help them. To this end, the varroa management concept developed by the Swiss Bee Health Service (SBHS) can be a valuable aid. All aide-memoires available at: www.ApiService.ch
Bee resistance to Varroa mites cannot be reduced to a single trait or an acronym like VSH or MNR. This article provides a clear and nuanced overview of the mechanisms currently under discussion, demonstrating why what truly matters in the apiary is a colony’s ability to sustainably curb the parasite’s growth and better weather the season.
The main environmental determinants of caste development in bees arise from larval nutrition: royal jelly stimulates the differentiation of larvae into queens, whereas bee bread leads to the development of worker bees. However, these determinants have not yet been fully characterized.
Apple and pear trees are among the main pome fruit trees in Switzerland. They are of great economic importance, and numerous varieties are cultivated. They are deciduous trees or shrubs. The flowers are hermaphroditic and are white to slightly pink with yellow anthers in the cultivated apple (M. domestica, originally M. sylvestris), whereas they are white (rarely pink) with dark red anthers in the cultivated pear (P. communis, originally P. pyraster).
Calming bees makes beekeeping easier and reduces stress for both the colony and the beekeeper. The summary below is based exclusively on the official checklist 4.10 from the Swiss Beekeeping Service (BGD/SSA) and refers to the complete PDF files.
This section compiles frequently asked questions about beekeeping, along with brief, practical answers that are directly applicable to the apiary. For each question, the goal is to get straight to the point: what to do, what to avoid, and what to check if things don’t go as planned. The aim is to provide simple, reliable, and easy-to-follow guidelines to help you make the right decisions at the right time.
Invasive plants are non-native plant species that have been introduced intentionally or accidentally by humans outside their area of origin and that reproduce and spread extensively, to the detriment of native species. Most often originating from other continents, they profoundly disrupt ecosystems and reduce local biological diversity.
(By Jean Riondet)
The question of feeding is recurring: in spring to stimulate colonies and to have strong populations at the time of the first nectar flows, then during the season in periods of dearth, and finally to ensure winter stores.
Any decision to feed colonies must be carefully considered, taking the season into account, have a specific purpose, and the ingredients must be well chosen so as to answer the questions properly: when ?, why ?, how ?
The list of recommended bee preparations provides an overview of products that are authorized and recommended for bee health. It serves as a guide for the selection and correct use of preparations in accordance with current regulations. The following information is based on the official document issued by the Swiss Bee Health Service.
Neonicotinoids are extremely ecotoxic pesticides. By infiltrating all environmental compartments, they indiscriminately kill both vertebrates and invertebrates. Despite their ban in 2018, traces of these substances persist in the environment at lethal concentrations.
Replacing frames is an essential part of good beekeeping practice and contributes significantly to the health of colonies. Regularly replacing old frames reduces health risks and improves brood and storage conditions. The following information is based on official guideline 4.4 from the Swiss Bee Health Service.
The expression may come as a surprise and calls for some clarification. “Insemination with a single male” refers to the insemination of a queen using the semen of a single drone, instead of the 12 to 20 males involved in natural mating or conventional instrumental insemination. This article presents this specific technique and its contexts of use. An overview of our current knowledge regarding the benefits and the present limitations of this technique!
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Pesticides do not necessarily remain where they are applied: residues can travel through air, dust, water, pollen, bee bread and wax into the hive environment. This article puts two recent analyses on exposure, regulation and bee health into context, with a particular focus on what the Swiss data suggest.
The Asian hornet is poorly named, since the vast majority of hornets (including our European hornet) originate from Asia. Scientists are more precise and refer to it as Vespa velutina nigrithorax. This Latin designation could be translated as “large wasp, covered with numerous short, silky hairs, with a black thorax”. It belongs to the hymenopteran insects (bearing 4 membranous wings that couple in flight via a series of small hamuli) Apocrita (with a narrow wasp waist). Its portrait is clearly less poetic...
The choice of location directly influences the health, development, and performance of bee colonies. Climatic conditions, nectar resources, and the immediate environment play a decisive role. The information below is based on the official checklist 4.9 of the Swiss Bee Health Service.
Monoecious deciduous tree or shrub. The flowers are unisexual, yellow-green, and arranged in racemes or panicles. The samara (fruit) consists of two seeds joined in a helical shape and is dispersed by wind. Species of importance for bees include: sycamore maple (A. pseudoplatanus), Norway maple (A. platanoides), and field maple (A. campestre).
Bees are remarkable animals. Did you know that they have existed for more than 65 million years (around 260 times longer than humans) and that they are the only insects in the world that produce a food consumed by humans without any processing.
Over the past year, articles dealing with the falsification of wax have frequently been published in the specialist press. The damage includes the collapse of combs, patchy brood patterns on newly built combs, and bees becoming trapped during emergence. The cause of these problems is the addition of stearin, paraffin, or other organic substances. Pesticide residues can also cause damage. As is known from experience with mothballs, there is likewise a risk that undesirable chemical additives in the wax may later appear in the honey.

A row of bee hives in a field of flowers with an orchard behind
© Catalin Petolea
Forming young colonies is a key tool for renewing the apiary, building reserves and managing colonies with greater flexibility. This article presents the main methods, their aims, limitations and key points of attention, especially regarding the queen, food supply, swarming tendency and varroa.
In Switzerland, as elsewhere, bees are disappearing, sometimes in a dramatic way. Without bees, it is not only honey that risks becoming scarce, but also fruits and vegetables. In an attempt to unravel the mystery of the disappearance of bees, Fred and Jamy travelled to the Vaucluse. Fred met with beekeepers and farmers, while Jamy set up his mini laboratory at the INRA in Avignon, right in the midst of hives used for research.
In Switzerland, beekeeping is considered a leisure activity. Today, around 17,500 beekeepers look after approximately 188,000 colonies, with an average of 11 colonies per person. In Germany, an operation with more than 30 colonies is regarded as a hobby activity, while a professional beekeeper manages more than 300 colonies.

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In Switzerland, ants are commonly found in or on beehives, but they are not generally considered a major pest of healthy colonies. This does not mean they are insignificant: they can exploit available resources, disrupt already weakened colonies, skew interpretations of natural varroa mite mortality, and play a role in the ecology of pathogens around the apiary.
The different bee races are determined using biometric traits (coloration, indices, etc.) as well as molecular markers (mitochondrial and nuclear). Defining races allows us to understand the biogeography of the honey bee.
Since antiquity, honey has been consumed for its sweetening properties and used for its therapeutic effects. Today, it has largely been supplanted in the diet by beet sugar. In medicine, however, its bactericidal and wound-healing effects on chronic wounds are being rediscovered. Beeswax is favoured by cosmetic manufacturers, while royal jelly and propolis are recognised in dietetics. The history of the use of honey and products of the beehive is fascinating.
Among the various goldenrod species with invasive characteristics are the late or giant goldenrod (Solidago gigantea) and the Canadian goldenrod (Solidago canadensis). It is rather difficult to distinguish between these two invasive species originating from North America, especially as they readily hybridise with one another.

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Below, you can observe in real time the variations in hive weight and the changes in colony temperature in different apiaries located in the plain and on the slope of the right bank of the Rhône.
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This article examines a question that is becoming increasingly relevant for apiaries in Switzerland and temperate Europe: how effective is winter treatment against varroa when colonies still have brood? It explains why the absence of capped brood remains a key condition for the effectiveness of oxalic acid — and why protecting winter bees starts well before December.
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The honey bee dance is not merely an instinctive reflex: its precision is also shaped by observing experienced foragers. Two recent studies shed light on the link between learning, vision and orientation — and help explain why visual landmarks, including around water sources, matter more in the apiary than one might think.
Varroa mites are the main parasitic disease affecting honey bees and, without rigorous control measures, pose a major threat to the survival of colonies. They are caused by the Varroa mite and weaken both the brood and adult bees. The following information is based on the official checklist 2.8 of the Swiss Bee Health Service.
When a colony is affected by an incurable disease or is severely weakened, it must be eliminated. This is an important measure within the framework of the control and prevention of epizootics and diseases. In the event of suspected notifiable disease, the apiary inspector must be informed before eliminating the colony.
The creation of nucleus colonies, alongside drone brood removal, is a central biotechnical measure to slow the development of varroa. Practical Guide 1.4 compares the methods described by the Swiss Bee Health Service (1.4.2–1.4.7) and supports decision-making. The basis is the official Practical Guide 1.4 “Overview of Methods for Nucleus Colony Formation” (V 2511).
In December, the apiary enters a quieter phase in which the undisturbed overwintering of the colonies takes priority. The beekeeper mainly observes from outside, checks the hives after severe weather, and carefully chooses the right window for the winter treatment against varroa without assuming broodlessness from the calendar alone. This quieter period is also useful for monitoring food stores, maintaining equipment, and preparing the next season.
Professor Martin Giufra works at the Center for Research on Animal Cognition in Toulouse, France. He is a specialist in neurobiology, with a particular focus on neurocognition in invertebrates. The research team he leads has investigated the remarkable learning abilities of the honey bee’s small brain.
Le traitement estival contre le varroa constitue un élément essentiel du concept de lutte contre le varroa. Il vise à réduire efficacement l’infestation après la récolte du miel afin de protéger l’élevage des abeilles d’hiver. L’aide-mémoire donne une vue d’ensemble des méthodes autorisées et de leurs conditions d’application. Les informations suivantes sont basées sur l’aide-mémoire officiel 1.2 du Service sanitaire apicole suisse.

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What happens inside a capped brood cell as a larva becomes an adult bee? This article explains honey bee metamorphosis and shows why this seemingly quiet stage is, biologically, one of intense transformation.
July is a pivotal month at the apiary. Between harvesting, the first summer treatment, establishing new colonies, and gradually preparing for winter, beekeepers must already be thinking well beyond this season’s honey. This month therefore calls for calm, well-organized decisions that are directly tied to the health of the colonies for fall and winter.
When a beekeeper looks at the head of his bees and sees the two large, immobile compound eyes positioned on either side of the head, as well as the three ocelli located on the forehead or vertex, he inevitably asks himself the question: with these two large eyes, can my bees see the same things as I do, or do they perceive the world differently? And why are there additional eyes on the head?
Dioecious tree or shrub with deciduous leaves. The male flowers have yellow anthers; the female flowers are inconspicuous, greenish, and develop at maturity into capsules rich in seeds.
The pavilion apiary dates back to the 19th century; its design was intended for storing baskets and hives with rear access. The apiary was meant to protect bees from wind, rain, and theft. The “Swiss” rear-access hive remains the most common management system in German-speaking Switzerland. Outside Switzerland, beekeeping is practiced mainly using multi-super hives.
European foulbrood is a bacterial disease of the brood in which the larvae usually die before capping. The official SSA/BGD checklist 2.2 describes typical signs such as patchy brood, yellowish to brown-black larvae, the match test, and a characteristic odor. It also specifies that this is a notifiable epizootic disease and that the apiary inspector must be called immediately.
The individual behavior of the honey bee is embedded in the overall management of the colony as a superorganism. Activity deep within the brood nest is now revealed thanks to the work of a research team that has developed a technology for digitally recording striking video footage. The intimacy of the colony is exposed down to the bottom of the comb cells: egg laying, larval hatching, care of open brood, and food storage appear in short videos, making it possible to visualize behaviors that had previously gone unnoticed.
Many animal species can be observed at apiaries, interacting directly or indirectly with bee colonies. This checklist presents the accompanying fauna, animals that are generally harmless and those that may cause problems, and describes the appropriate way to deal with these situations. The following information is based on the official checklist 3.4 of the Swiss Bee Health Service.
There are many methods of queen rearing. All of them are based on the same principles. Every beekeeper knows that, outside the swarming period, a colony with a queen will not accept the rearing of additional queens without specific precautions (in general, the presence of the queen prevents or destroys any queen rearing). Queen-rearing techniques consist in the practical implementation of these “precautions”.
Bees do not close their eyes throughout their lives. Nevertheless, their activities are influenced by an intracerebral protein (pigment-dispersing factor, PDF), which enables them, among other things, to orient themselves in relation to the position of the sun.
For honey bees, willow is one of the first food sources at the end of winter. Beekeepers time their spring inspection according to the flowering of the catkins of the male plant.
Feeding is an essential part of colony management and helps to compensate for periods of food shortage while ensuring the development and wintering capacity of colonies. The timing, quantity, and type of food must be adapted to the strength of the colony and the season. The information below is based on the official checklist 4.2 of the Bee Health Service (SSA).

Melezitose | melicitose | nonreducing | trisaccharide | sugar | honey | bee | problem | crystallized | granulated | comb | wax | bee bread | open | agriculture | animal | apiary | bee | beehive | beekeeping | beeswax | biology | busy | cell | close | colo
Melezitose is not a defect of forest honey, but a natural sugar found in certain honeydew flows. It becomes problematic mainly when honey crystallizes already in the combs or when such stores remain in the brood chamber over winter. This article explains its origin, mechanisms, and practical implications for harvest and overwintering.
In March, the apiary comes back to life: the queen gradually increases her egg-laying, foragers fly as soon as temperatures allow, and food consumption rises to support brood development. This is a pivotal period in which the beekeeper must observe each colony calmly, as a lack of resources or an undetected disease can compromise recovery after winter.
The wax moth is not considered a disease. In nature, wax moths play an important role by destroying old, abandoned combs, which are potential sources of pathogens. Attracted by odours, wax moth adults enter hives and lay their eggs there. The resulting larvae feed on pollen residues and on cocoon remnants that remain at the bottom of the cells.
While the bee amazes us with its highly social behaviour, its anatomy and multifunctional organs are truly awe-inspiring.
The bee is equipped with two antennae attached to the upper part of its head, composed of three main segments: the scape, the pedicel, and the flagellum. The antennae are in constant motion thanks to four muscles located in the head capsule that enable rotation of the scape; the latter contains two groups of muscles responsible for the movements of the flagellum and the pedicel. The motor neurons involved in activating these muscles (nine for the scape, six for the flagellum) have their cell bodies located in the dorsal lobe, which constitutes the motor and mechanosensory system of the antennae. The antennae thus serve as the bee’s central organs for perceiving its environment and, through thousands of sensilla, transmit an enormous amount of information linking it to the external world. The antennae therefore support the senses of smell, taste, touch, hearing, and the perception of electric fields.
Pollinators use precise cues to identify flowers rich in pollen or nectar: colour, scent … Researchers have discovered that bees are also sensitive to temperature variations between different parts of a flower. These “thermal patterns” enable them to locate the flowers richest in food.
The establishment of a flying nucleus colony (a nucleus formed at midday from a collective brood nucleus with an additional rearing unit) serves to deliberately supply young colonies with forager bees during the day and to promote their development. The following summary is based exclusively on the official Practical Guide 1.4.5 (BGD/SSA) and refers to the original source.
Asian bees have developed an extraordinary technique to defend themselves against attacks by giant hornets. They cover their hive with animal droppings.
Like any living organism, the honey bee can become ill. The beekeeper must remain vigilant, as a disease can have serious consequences, especially if it is a so-called “notifiable contagious disease” (MRC). A beekeeper who has been practising this activity for many years will immediately detect even the slightest anomaly in the hives. For a beginner, however, it is often very difficult to identify a disease; this is why they must be much more observant, considerably more inquisitive, and always very attentive to what is happening on the landing board.
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Learning, remembering, generalising: the honey bee performs surprisingly complex tasks with a tiny brain. Martin Giurfa’s review shows what neuroscience experiments reveal about these abilities — while also reminding us why laboratory findings need to be interpreted with care.
Das Bannwabenverfahren ist eine biotechnische Methode zur Varroabekämpfung, bei der die Eiablage der Königin gezielt auf bestimmte Brutwaben gelenkt wird. Dadurch kann die Brut und damit ein grosser Teil der Varroamilben kontrolliert entfernt werden. Die folgenden Hinweise basieren auf dem offiziellen Merkblatt 1.6.2 des Schweizerischen Bienengesundheitsdienstes.
Although they are few in number, wild or feral colonies of A. mellifera mellifera do indeed exist in nature and are resistant to Varroa due to their small size, their tendency to swarm, the low density of the habitat, and probably through natural selection. Studies also show that these strains of black bees are pure, without hybridization. Consequently, researchers argue for the development of a conservation policy for these bee populations, because they constitute a wild taxon that is probably in decline, but also an interesting genetic reservoir from a beekeeping perspective.
Buddleia (scientific name: Buddleja davidii) is a shrub also known as the “butterfly bush”. Despite its appealing name, it is an invasive plant whose spread should be strictly limited.
Preventing swarming does not consist simply in giving the queen more room or removing queen cells. Swarming is a gradual transformation of the colony: depending on when the beekeeper intervenes, one and the same operation may act as prevention, as a modification of preparation already under way, or only as a means of avoiding the loss of the swarm.
Recognising bee diseases and knowing effective control and prevention methods are sine qua non conditions for ensuring bee health and for good beekeeping practice. Diseases spread not only very rapidly within hives because of physical contact between worker bees and trophallaxis (the exchange of food between bees), but also between hives. Because bees can fly over long distances, rob neighbouring colonies, or drift into them, the risk that a large number of colonies and apiaries will be affected by a disease or an epizootic is significant. When one also considers the movement of hives by beekeepers and the high density of apiaries in Switzerland, the prevention of epizootics and diseases becomes even more important in bees than in other livestock animals, which can be quarantined more easily.
Bee paralysis is a viral disease that mainly affects adult bees and can lead to behavioral disorders and increased losses. Official memo 2.10 from the Swiss Bee Health Service describes the symptoms, contributing factors, and what to do if you suspect your bees are affected.
Is it because it provides honey and wax? Or because its sophisticated organisation strangely echoes human societies? Or perhaps because of a character that is at once fierce and disciplined, making its domestication uncertain, or because of the way it positions itself at the confluence of the plant and animal worlds. Protean and tinged with mystery, the fascination exerted by the bee has endured for millennia, albeit with periods of interruption.
The brood nucleus colony is an established beekeeping method for creating new colonies based on brood combs from existing colonies. It serves colony multiplication, swarm prevention, and can contribute to reducing varroa pressure. The following information is based on the official Practical Guide 1.4.4 of the Swiss Bee Health Service.
by SEAN BAILLY
Aurore Avarguès-Weber from the University of Toulouse and colleagues from Melbourne, Australia, under the direction of Adrian Dyer, have continued to explore the mathematical abilities of bees. They have now shown that bees are capable of addition
and subtraction.
Bees play an essential role in our ecosystem because of their primary function, pollination. This activity enables plants to reproduce. Alongside wasps and butterflies, bees are among the most efficient pollinating insects. However, they remain a species that is severely threatened by various factors.
Waste control makes it possible to draw numerous conclusions about the health status of a colony. Frequent checks do not disturb the colony. By examining the bottom boards, anomalies and irregularities can be identified, providing indications for future interventions to be carried out on the colony.
This contagious disease of the honey bee is caused by SBV, the abbreviation of its English name Sacbrood Bee Virus. It is present worldwide and generally affects capped brood, leading to more or less significant mortality of prepupae, which may result in colony weakening.
This disease is characterised by the typical sac-like appearance of the prepupae killed by the virus. Infected adult bees show no symptoms (they are asymptomatic carriers) but constitute reservoirs of the virus.
Perennial herbaceous plants with toothed leaves arranged in a basal rosette. The robust stems bear large inflorescences composed of numerous bisexual flowers, which may be yellow, orange or blue. The spherical infructescence consists of seeds that are dispersed by the wind thanks to their umbrella-shaped pappus.
In Switzerland, for example, the common dandelion (Taraxacum officinale), chicory (Cichorium intybus), as well as various species of hawk’s-beard (Crepis sp.) and hawkweed (Hieracium sp.) belong to the so-called “dandelion forms” (whose pollen is difficult to distinguish).
by Yves Martignoni and Julien Balet
What do breeding instructors do? What is breeding? Is it something within my reach? These are some of the questions we will attempt to address in this article.
In nature, bees rear queens in spring during swarming in order to multiply colonies. They also rear queens in other situations, for example when the queen no longer performs satisfactorily after several years of egg-laying, or when the queen is accidentally lost. In these particular cases, queen rearing is not associated with swarming.
Yellow, orange, red, blue, grey or almost black: the pellets brought back to the hive offer a glimpse of the resources being exploited. Their colour can guide observation, but it is not sufficient either to identify a plant with certainty or to judge the nutritional quality of the pollen. The table proposed here should therefore be used as a field guide, not as an identification key.
The bee is an insect belonging to the order Hymenoptera (together with wasps, ants, etc.). Hymenopterans are equipped with two pairs of wings, three pairs of legs, and articulated appendages. Their body is composed of three distinct parts: the head, the thorax, and the abdomen.
Phacelia, Phacelia tanacetifolia, is an annual herbaceous plant reaching a height of between 50 and 120 cm. It originates from California and Mexico and was introduced into Europe in the 19th century. It occurs near dwellings and on wasteland, where it can even become invasive; however, it is mainly cultivated as a green manure, as a forage crop, or as an intermediate crop.

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Crowded hives are not only a matter of apiary layout: they may also increase drifting and the spread of varroa between colonies. The study by Seeley and Smith shows how spacing, robbing and reinvasion can influence overwintering success. For beekeepers, the practical message is clear: good apiary organisation can reduce risk, but it never replaces rigorous varroa monitoring and control.
The creation of collective brood nuclei serves colony multiplication, supplementary feeding, or the rehabilitation of weakened stocks. This section summarises the key points of the official Practical Guide 1.4.4.1 and refers to the complete original source.
The introduction of a queen is an essential operation in beekeeping to replace an existing queen, complete a nucleus colony, or guide selection. This summary is based exclusively on the official checklist 4.5.2 of the Swiss Beekeeping Service (BGD/SSA).

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Is a bee that is staggering necessarily “drunk”? Experiments show that ethanol can disrupt its behavior, but the disturbances observed in the apiary may have many other causes. This article reviews the demonstrated effects of alcohol and the observations needed to examine its role under natural conditions.
by Sophie Bécherel
Dopamine is a neurotransmitter involved in the cerebral reward system and the sensation of pleasure in mammals. While in humans dopamine is a key element in the mechanism of drug addiction, in bees this neurotransmitter is responsible for the departure of foragers from the hive, not only to satisfy their individual desire to feed, but also to search for food in order to meet a social need and provide the colony with the nutritional inputs it requires.
Treatment with formic acid strips (Formicpro®) is an approved method for controlling the varroa mite during the brood-rearing phase of the honey bee colony. It enables controlled release of the active substance over a defined period. The following information is based on the official Practical Guide 1.2.5 of the Swiss Bee Health Service.
May is an intense period in the apiary. Colonies gain strength quickly, honey supers need to be added at the right time, swarming pressure often increases, and sound colony health management remains essential. Depending on the region, altitude, and weather, this month may already allow for a first spring honey harvest.
by AURORE AVARGUÈS-WEBER
Despite having a tiny brain with 100,000 times fewer neurons than ours, bees possess remarkable cognitive abilities. These hymenopterans can count, master concepts, reason by categories… and are even faster than great apes in certain tasks!
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Winter losses do not start only in winter. This article explains why the decisive phase begins in late summer, when long-lived winter bees are produced and their quality is shaped by Varroa, nutrition, brood dynamics, and colony stress.
In September, the apiary enters a critical phase: the health of the hives, their food stores, the quality of the queens, and the culling of colonies now take precedence over honey production. This month lays the groundwork for wintering: effective varroa treatment, replenishing food stores in a timely manner, and not artificially prolonging the life of colonies with no future are decisions that will directly impact the following spring.
Tropilaelaps is expanding beyond its original Asian range and spreading westwards, bringing it increasingly into focus for European beekeeping. This article reviews what is actually known about its biology, population growth and detection, and what this may mean for beekeeping in Switzerland.
April marks a decisive phase in the development of colonies. After winter, the population grows rapidly and activity in the apiary intensifies. Space management, observing the first signs of swarming, and health monitoring become priorities. Interventions must remain measured and be based above all on observation of the actual dynamics of the colony and the environment.
This study shows how two small experimental colonies coordinated ventilation, waste removal, and water collection when their brood nest was artificially heated. It also highlights temporary water storage in the crop of some workers and, to a lesser extent, in cells, without demonstrating the existence of a long-term reserve.
Melliferous plants, apicultural plants, mellitophilous plants… all these terms refer to plants that are of interest to bees. But what do they really mean, and which term is the most appropriate?
Sea buckthorn, whose scientific name is Hippophae rhamnoides, is an evergreen plant belonging to the botanical family Elaeagnaceae. This hardy and highly ornamental shrub, which can reach a height of up to 4 metres, has a densely branched stem with thorny shoots on which small, fleshy, orange-coloured berries develop, resulting from the development of the floral receptacle after fertilisation.
Winter is undoubtedly the most demanding period for bees. However, nature has provided so-called “winter bees” with characteristics that enable them to withstand the rigours of the cold season. Winter bees have larger fat reserves than summer bees, they possess a more developed hair covering that provides natural insulation, and the contraction of the thoracic muscles generates a significantly greater release of heat than in summer bees. Their constitution, together with their behaviour, allows them to live longer and to cope with the harsh conditions of winter.
It is a contagious disease of the honey bee caused by a virus known as CBPV, an abbreviation of its English name Chronic Bee Paralysis Virus (chronic bee paralysis virus).
Nutrition appears to play a particularly important role in the health and immune defences of bees. They must have continuous access to nectar and pollen. However, this requirement is difficult to meet in modern agricultural landscapes. Periods without nectar flow during the most intensive phase of brood rearing slow colony growth and result in increased susceptibility to disease.

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Honey production involves more than just harvesting and extraction; it also falls within a legal framework that encompasses primary production, product regulations, and traceability. This reference guide provides Swiss beekeepers with a practical overview of the key points they need to know, which are then presented in a structured format in a summary table.
This protocol sheet facilitates the structured documentation of suspected bee poisoning and provides precise instructions for taking and sending samples. It is intended for beekeepers and competent persons to support coordinated clarification. The content is based on the official checklist 3.1.1 of the Swiss Bee Health Service.
Chalkbrood is a fungal disease that affects worker and drone brood. Contamination by spores occurs through larval feeding. The fungus germinates in the larval gut and spreads throughout the body in the form of filaments. The disease occurs mainly in weak colonies; it is promoted by sharp drops in temperature and high humidity. It may affect individual colonies or, under unfavourable weather conditions (cold, humidity), entire apiaries in the form of an epidemic. A site that is regularly and heavily affected by chalkbrood is considered unsuitable; hives should therefore be relocated to a sunnier location. Severe infestation can kill colonies.
Within the colony, the role of the drone is often poorly understood or even regarded as secondary. Yet males play a fundamental role in the transmission of genetic diversity. Gathered in so-called drone congregation areas comprising 1,000–15,000 individuals, they originate from many different colonies, sometimes located 10–15 km away. During the so-called drone flight, they mate with queens in succession (often up to 20 consecutive matings), thereby ensuring genetic mixing and preventing inbreeding.
Spray treatment is a biotechnical method for varroa control, primarily applied to broodless or nearly broodless honey bee colonies. It is based on the direct wetting of the bees with an approved solution and requires precise and careful handling. The following information is based on the official Practical Guide 1.3.1 of the Swiss Bee Health Service.
The quality of syrups used for winter feeding of bees is very often at the centre of lively discussions.
What should be the main qualities of a syrup:
Natural comb construction involves building without embossed wax foundation, giving bees the freedom to build their combs according to their needs. This fact sheet outlines the conditions, advantages, and limitations of this beekeeping practice. The following information is based on the official fact sheet 4.4.3 from the Swiss Bee Health Service.
Horizontal or creeping cotoneaster (Cotoneaster horizontalis) is a shrub native to China that was introduced to Europe at the end of the 19th century. For a long time, it remained confined to parks and gardens, where it is appreciated for forming a dense ground cover that blankets rock gardens and stabilises slopes.
The dandelion, scientifically known as Taraxacum officinale and commonly referred to as “dandelion,” is a flowering plant belonging to the Asteraceae family. It is a perennial herbaceous plant (living for several years) that is widely distributed across a wide range of habitats, particularly on nutrient-rich soils, in lowlands as well as in mountainous regions up to elevations of about 2,500 m. Flowering begins in April and reaches its peak in May, but continues at a lower intensity into autumn, especially during dry and warm summers.
The article explains how the colony's internal biological mechanisms provide a better understanding of seasonal variations in development, the importance of harvests during favorable honey flows, the transition from summer bees to winter bees, and certain phenomena of decline.
The scientific work presented highlights the central role of vitellogenin, brood pheromones, and protein nutrition in the social organization, survival, and ability of bees to adapt to their environment.
Translated with DeepL.com (free version)
The wash method (also referred to as the “wash tower”/“washing procedure”) is used to wet bees with a liquid in order to dislodge and quantify Varroa mites from brood chambers and from the bodies of the bees. It is used for diagnostic purposes and does not replace treatment. The following summary is based on the official Practical Guide 1.5.3 of the Swiss Bee Health Service.
The bee’s mini brain, which contains about 10⁶ cells (10¹¹ in humans), enables it not only to manage stereotyped behaviors such as foraging, but also—thanks to its plasticity—to adapt and allow the insect to respond to new problems through often complex learning. The bee’s brain is capable of providing “intelligent” solutions to a wide range of ecological or other problems, as is the case in vertebrates and in humans.
Communication in bees is highly elaborate and has been the subject of numerous studies. There is, of course, the well-known “waggle dance” or “figure-eight dance,” but what is less well known is that this dance serves solely to indicate the location of a food source. While dancing, the bee releases chemical messengers that recruit other foragers and inform them about the type and richness of the source. Communication therefore indeed relies on the exchange of chemical substances known as pheromones.
The natural swarm is the natural form of colony reproduction and occurs primarily in spring and early summer. It represents both an opportunity for colony multiplication and a challenge for apiary management. The following information is based on the official Practical Guide 1.4.6 of the Swiss Bee Health Service.
Chemical control of harmful organisms has been known for millennia. The application of sulfur, lead, or arsenic was already practiced in ancient Greece. Closer to our time, the insecticidal properties of tobacco (nicotine) were known as early as the 17th century. In the 19th century, chemistry made major advances, and new copper sulfate–based compounds made it possible to control fungal diseases of grapevines. In the 20th century, mercury salts began to be used for seed treatment. Because of their toxicity, these substances were quickly banned.
Forming a summer nucleus from bees in the supers may seem straightforward at harvest time. The method can work, but late in the season its success depends mainly on how quickly the colony gets established, on the resources available, and on the nucleus's capacity to build a viable autumn population in time.
By Claude Pfefferlé and Serge Imboden
Long regarded as a model of uninterrupted activity, the honey bee nevertheless continues to surprise researchers. Recent studies have revealed that it sleeps, and that this sleep plays an essential role in regulating its memory, physiology, and group cohesion. Understanding how and why bees sleep is to open a window onto the health of the hive—and onto the balance of life itself.
A nightmare for some, an opportunity for others, an incredible hope for visionaries, the wax moth is an insect that intrigues all those interested in its evolution and its perfect adaptation to the hive and its superorganism. There are therefore three ways of viewing the wax moth: the beekeeper fears the damage it causes in hives; the entomologist marvels at the undertaker role played by the insect; researchers, for their part, focus on its highly ecological ability to digest highly polluting plastic. So what is this curious moth?
Can honey bees survive in the long term without beekeepers? Free-living colonies also exist in Europe, but the high losses observed in Swiss studies call for caution. This article examines survival mechanisms and natural processes of resistance to Varroa mites in order to identify lessons that can be applied to beekeeping practice.
Migratory beekeeping allows colonies to benefit from different nectar flows throughout the year, but it requires thorough preparation and adherence to precise rules. Official Practical Guide 4.9.1 of the Swiss Bee Health Service describes the conditions, organisational requirements, and principles to be observed when moving colonies.
American foulbrood is a bacterial disease. Under optimal conditions, the bacteria multiply by cell division. When conditions become unfavourable, spores are formed that remain infectious for decades. These highly infectious spores are distributed within the colony in the brood nest through the cleaning and maintenance activities of worker bees. Infection by spores occurs via the royal jelly on which larvae feed during the first 48 hours of their life. Adult bees are not infected but can act as carriers of the pathogen. A completely dried larva forming a crust may contain more than 2 billion spores.
Targeted drone brood removal is a practical measure for reducing the varroa population. It uses the different brood characteristics of drones to remove mites from the brood nest and interrupt their reproductive cycle. The following summary is based on the official Practical Guide 1.4.1 of the Swiss Bee Health Service.
Deciduous trees and shrubs with hermaphroditic white to slightly pink flowers that develop into edible fruits of various colours. In Switzerland, the wild cherry (P. avium), plum (P. domestica), sour cherry (P. cerasus), apricot (P. armeniaca), and blackthorn (P. spinosa) are commonly found.
The art of economy (Janine Kevits)
Winter represents a formidable challenge for fauna, as it must cope both with cold temperatures and with food scarcity. Some insects have “chosen” to avoid it by migrating to warmer regions; this is the case, for example, of the painted lady butterfly. Others concentrate their chances of survival on a few individuals—reproductives that are abundantly nourished during the favorable season and whose task is to found a new colony on their own the following spring; this is the strategy of wasps, hornets, and other solitary bees. The honey bee, by contrast, has found a different path: it is the powerful organization of the colonies it forms that enables it to meet this challenge, by implementing two means that are entirely original in the insect world—on the one hand, the storage of reserves, and on the other, the reorganization of the colony to form the winter cluster, a system characterized by the absence of brood and by modes of functioning that differ fundamentally from those of the summer colony.
Vaporisation/sublimation of oxalic acid is a winter treatment recommended in Switzerland against varroa, provided that colonies are broodless or contain only open brood. Vaporisation is carried out using a dedicated sublimator/vaporiser device; correct application is essential for both efficacy and safety. The following summary is based exclusively on the official Practical Guide 1.3.3 (V 2511) of the SSA / BGD.
Varroa is known to suck the haemolymph of bees. Well known, but apparently (largely) incorrect.
Researchers in the United States were intrigued by the magnitude of varroa’s impact on bee health in relation to the relatively small amount of haemolymph it extracts. Since insect haemolymph is comparatively poorer in nutrients than mammalian blood, they questioned how the parasite could develop on such a resource.
Observation at the hive entrance makes it possible to draw numerous conclusions about the health status of a bee colony. Especially in spring, it is important to assess colonies in this way. Anomalies indicate the aspects that require particular attention during the next colony inspection or when an additional examination becomes necessary. Observation at the hive entrance also helps to avoid interventions within the colony under unfavorable weather conditions.
Varroosis promotes drifting, a mechanism explained by the fact that bees from heavily infested colonies lose the specificity of their cuticular chemical profile as well as the accuracy of their orientation. What can we learn from wild colonies?
The disease is caused by a virus. The pathogen can multiply both in adult bees and in the brood. In contrast to elongated larvae or prepupae, which may become diseased and die, no clinical symptoms are visible in infected adult bees, but their lifespan is reduced.
Gilbert Duruz is a beekeeper in Switzerland, in the upper Rhône Valley. He is a beekeeping trainer, queen breeder, tester for the Swiss Romand selection programme of the Carnica bee, and president of the Association Suisse Romande des Éleveurs de Reines (ARAE). He presents his breeding and renewal method under continental and mountain conditions.
Himalayan balsam or glandular balsam (Impatiens glandulifera), of Asian origin, is a typical example of a plant that divides beekeepers, gardeners, and ecologists: highly nectar-producing and very ornamental, it nevertheless poses an ecological threat, as its invasive nature has a substantial impact on local biodiversity.
Asters are perennial herbaceous plants belonging to the Asteraceae (or Compositae) family. Their name derives from the Greek aster, meaning star or flower.
They are ornamental plants widely cultivated in gardens, with more than 250 species. Most originate from North America, some were introduced from Asia, while others are native to Europe.
The merging of bee colonies is a method used to strengthen weak colonies or compensate for losses. It must be carried out in a manner appropriate to the bees, taking into account the strength of the colonies, the season, and weather conditions. The summary below is based exclusively on the official checklist 4.7.1 of the Swiss Beekeeping Service (BGD/SSA).
January generally corresponds to the core of the overwintering period in Switzerland. The colony lives at a reduced pace in a cluster: it maintains a sufficient internal temperature to protect the queen, and if brood is present (which varies depending on altitude and climate), the central area is heated more intensely. Honey consumption remains moderate but continuous. Typical risks of the month include “silent” starvation (insufficient or inaccessible reserves), humidity, and disturbances. The beekeeper therefore acts mainly through external observation, targeted securing of reserves, hygiene, and preparation of equipment, while avoiding any intervention. The watchword is: “Do not disturb!”
The organs of the digestive system of the honey bee enable the assimilation of food. Certain glands are associated with the digestive system while also fulfilling peripheral functions, such as the production of nutritive substances or supporting food assimilation.
For millions of years, bees have ensured the survival of their species through swarming. Swarming is a natural process whose main objective is to create new, healthy and genetically diverse colonies (the males that will mate with the virgin queen in a new territory have a different genetic background). Beekeepers, however, want colonies that develop well and become strong without swarming. A colony with swarming intent (swarming fever) no longer builds comb and collects little nectar. The summer harvest is often lost. How can this swarming fever be suppressed? A new method developed by the Sion Beekeeping Association (www.ApiSion.ch) appears promising.
Deciduous, monoecious, wind-pollinated shrub or tree. The pendulous male inflorescences consist of numerous small individual flowers. The small female flowers are enclosed in a bud from which only the red stigmas protrude and develop into oval, woody hazelnuts.
Good beekeeping practice: Every beekeeper should try to rear a few queens in order to maintain and improve the quality of their stock. In this short course, I will attempt to present a queen-rearing technique that incorporates certain tips used by royal jelly producers. This method requires only minimal investment, and no queen is lost.
The Nassenheider evaporator is an approved system for the controlled application of formic acid in varroa control. Correct handling as well as consideration of temperature, dosage, and safety measures are essential for the efficacy and tolerability of the treatment. The following information is based on the official Practical Guide 1.2.2 of the Swiss Bee Health Service.
Swarming is not a management failure, but the honey bee colony’s natural mode of reproduction by fission. This reference article explains how the colony prepares over days to weeks, how the swarm and mother colony are formed, and what this means for beekeeping practice.
Throughout the year, the bee colony is constantly changing: it grows, reaches its peak, and then gradually prepares for winter. At the same time, Varroa destructor follows its own dynamic, often less visible but crucial to the health of the hive. Understanding these two processes helps us better understand what is happening in the apiary and why certain times of the year are particularly critical. This article provides a clear overview of the possible trajectories throughout the season.
The idea that an animal or plant is merely the product of the genes inherited from its parents is increasingly being questioned. Until now, it was known that the environment can influence the appearance or behavior of living beings (the phenotype) to a considerable extent. However, it has been shown that in some cases, environmental influences can be passed on to subsequent generations without changing the genetic information itself. The entirety of the mechanisms that control this heritable component influenced by environmental factors is referred to as “epigenetics.”
by ELIZABETH TIBBETTS AND ADRIAN DYER
Recognizing the facial features of conspecifics does not require a brain as complex as one might imagine: some insects, notably bees, are remarkably adept at this.
Faced with the current context—polluted environments, pesticides, climate change, varroosis, etc.—beekeeping is confronted with multiple challenges. However, this context does not explain everything. In order to act effectively, it appears necessary to address the “health aspect” of bee colonies in a holistic manner. This approach assumes that the biological functioning of bees is similar to that of other living organisms and that all these factors share the characteristic of being influenced by human actions. Such reflection should guide beekeeping interventions in a way that respects the health of the bees.

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Beeswax is not just a building material: it can also preserve the chemical memory of treatments and of the apiary environment. This article explains why certain residues accumulate in wax, what this may mean for colony health, and why regular comb renewal remains an essential element of good beekeeping practice.
Unlike social insects, which include our honey bee Apis mellifera, wild or solitary bees have a different way of life. They do not share their nests, do not cooperate in brood rearing, show no overlap of generations, do not store honey, work independently, are not aggressive, and often lack a sting.
This official factsheet describes the use of the FAM dispenser with Formivar 70% and focuses on protective measures, required equipment, key points, and a clearly structured procedure. It specifically emphasizes that formic acid (Formivar) is highly corrosive and must be handled with great care. For details and the complete instructions, the factsheet is considered the authoritative reference.
To rear drones or not to rear drones – that is the question posed to our bees with the return of the favourable season. Within a colony, males are indeed a luxury: they are far more costly to rear than workers. And the expense does not end at emergence: as adults, drones largely remain dependent on their sisters. In addition, they contribute nothing to foraging activities and very little to hive maintenance. Yet this luxury is necessary: the sole function of males, the transmission of genes, lies at the very heart of the biological meaning of life. Maximising the chances of reproduction while preserving those of survival – it is this subtle balance, to which both the queen and her workers contribute, that we propose to explore today.
Some acaricides used in alternative control strategies against Varroa, such as formic acid or essential oils, do not always provide sufficient efficacy. As complementary measures, we recommend the removal of drone brood or the establishment of young colonies in spring. These interventions aim to slow the development of Varroa populations and thereby reduce infestation pressure. They have the advantage of being applicable during the main beekeeping season, whereas the use of chemotherapy would entail significant risks of contaminating honey harvests.
The activity of insects, which are cold-blooded animals, is determined by ambient temperature. Their muscles require a minimum level of heat; if this threshold is not met, all activity ceases. To avoid this potentially lethal phenomenon in winter, insects have developed resistance strategies, most commonly through hibernation.
When a bee changes its role in the hive, its DNA adapts
Bees each perform a specific but flexible role within the hive. Older individuals may, if necessary, take over tasks usually carried out by younger bees. Until now, it was unknown that such behavioural changes are accompanied by adaptations at the DNA level. Each role is in fact associated with its own distinct methylation pattern.
The garden aster is a perennial herbaceous plant reaching 90–150 cm in height and flowering from August to November. Native to North America, Aster novi-belgii was introduced to Europe in the 18th century as an ornamental plant. Having escaped cultivation, it now grows spontaneously in many places, preferably on rather calcareous soils: in gardens, meadows and moist woodlands.
Nature ensures colony multiplication through natural swarming, but the beekeeper desires colonies that develop well and become strong without swarming. A colony in the swarming impulse no longer engages in comb-building activity and collects little nectar. Swarms are frequently lost, particularly for the beekeeper located far from their apiary. Moreover, a parent colony that has swarmed requires special attention and care. What can the beekeeper do?

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This study shows that the royal jelly produced by nurse bees is not only used to feed the brood and the queen, but is also distributed to a significant portion of the adult bees. It thus sheds light on the central role of nurse bees in the redistribution of proteins within the colony. Pollen remains in the background as the primary resource, but it is the circulation of this glandular food that the study directly highlights.
The powdered sugar method is a non-destructive diagnostic technique for estimating varroa mite infestation in honey bees. It provides an estimate of the mite load by dislodging mites from the bees’ bodies using powdered sugar, allowing them to be counted. The following summary is based on the official Practical Guide 1.5.2 of the Swiss Bee Health Service.
The black locust or false acacia, known by its scientific name Robinia pseudoacacia, is an exotic plant that was introduced from North America in the 17th century. This tree, which can reach up to 25 metres in height and live for as long as 300 years, has spread easily throughout Switzerland, as it adapts to all types of soil and is resistant to pollution, to the point of sometimes being considered invasive; however, it is not found above 800 metres in altitude.
Herbaceous annual or biennial plant bearing racemose inflorescences with yellow flowers and long siliques (fruits). Oilseed rape is an important agricultural crop, cultivated as fodder or for its oil-rich seeds used in industry and food production.
It may happen, for example during the first spring inspection, that one encounters a moribund colony with no food reserves, with frames showing scattered brood of the male type … What has happened?
Figure 1: Drone-laying colony: very small population, scattered drone brood, no worker brood, low food reserves, no visible queen, deformed combs, drone cells.
by R. Prasad
The defense of a society often requires certain specialized members to coordinate in order to repel a threat at the risk of their lives. This is particularly true for honey bees, which defend the hive and may sacrifice their lives when stinging. At the core of this cooperative defensive response lies the sting alarm pheromone, whose main component is isoamyl acetate (IAA).